Aclista insolita Nixon, 1957

Macek, Jan, 2005, Revision of the Central European species of Aclista (Diapriidae, Hymenoptera). Part II. Aclista insolita Nixon, 1957 Aclista dubia Kieffer, 1909 and similar species, Acta Entomologica Musei Nationalis Pragae 45, pp. 183-197 : 84-85

Aclista insolita Nixon, 1957

( Figs. 1 , 8 , 15 , 22 , 29, 36 )

Aclista insolita Nixon, 1957: 66 , 80, J, ♀.

Acoretus insolitus: HELLÉN (1964) : 20, 23.

Aclista insolita: WALL (1967) : 157, 161.

Type locality. England, Somerset, Porlock district.

Type material. HOLOTYPE: ♀, ‘ England, Somerset, Porlock distr. , 21.v.1934, Perkins lgt., G.E.J Nixon det.’ ( BMNH) . PARATYPES: JJ ♀♀, ‘ Sweden, Skåne’ ( BMNH).

Additional material examined. CZECH REPUBLIC: BOHEMIA bor., Kadaň (5645), 3.vi.1982, 12 spec. ; BOHEMIA centr., Kublov (6049), 14.v.1993, 24 spec. ; BOHEMIA mer., Olešník (6852), 11.vi.1982, 22 spec. ; BOHEMIA occ., Zbiroh (6148), 3.vi.1985, 11 spec. ; BOHEMIA or., Orlické hory Mts., NR Trčkov (5764), 25.v.1994, 43 spec., J. Ježek et J. Hájek lgt. MORAVIA mer., Tvrdonice (7267), 15.v.1984, 13 spec. SLOVAKIA mer.: Silica (7489), 15.v.1986, 14 spec. ; Gombasek (7588), 17.v.1986, 11 spec. POLAND mer.: Ochotnica Gorna , 8.vi.1989, 1 spec ; all J. Macek lgt. & det. unless stated otherwise.

Diagnosis. Head in front view triangular; mandibles short and feeble, shortly overlapping at tips; mouth aperture narrow; toruli separated by deep cleft; eyes shorter than genae; both scape and median flagellomeres thickened in middle; epomia strongly developed; notauli deep, diverging posteriorly; radial cell longer than marginalis; pygidium large, longer than prepygidium; prepygidium with narrow, partly telescoped segments; aedeagus narrowed apically.

Variability. Female specimens with extensive variation range in following characters: colour brown or black; appendages pale or dark; flagellomeres rectangular, quadrate or transverse; face smooth or finely rugose; subantennal suture distinct or indistinct, becoming confluent with subantennal rugosity; pronotal shoulders sharply prominent or angular; radial cell long or short and narrow or wide (both attributes combined); radial vein straight or arcuate; plicae of propodeum parallel or slightly converging and projecting or not projecting posteriorly (both attributes combined); petiole stout or slender and smoothly ribbed or finely rugose (both attributes combined); gaster stout or slender. Males without any such variation.

Differential diagnosis. Aclista insolita is easily recognised from other Aclista by having both the scape and the flagellum distinctly thickened in the middle (only in females), short and feeble mandibles, a deep cleft between the toruli and indistinctly foveate facial grooves confluent with the subantennal rugosity (the latter three characters in both sexes).

Bionomy. Hosts unknown; a vernal species with flight period from April to June.

Distribution. The species is known from England ( NIXON 1957), Switzerland ( WALL 1967), the Czech Republic ( MACEK 1989), Finland (HELLÉN 1964), Sweden ( NIXON 1957) and European Russia ( KOZLOV 1978). New for Poland and Slovakia.

Figures

Figs. 1-7. Head, frontal view (female). 1 – Aclista insolita Nixon, 1957; 2 – A. vernalis sp. nov.; 3 – A. dubia Kieffer, 1909; 4 – A. longistilus Kieffer, 1909; 5 – A. marginalis Kieffer, 1909; 6 – A. elegantula sp. nov.; 7 – A. subtilis sp. nov. Scale = 1 mm.

Figs. 8-13. Head, dorsal view (female). 8 – Aclista insolita Nixon, 1957; 9 – A. vernalis sp. nov.; 10 – A. dubia Kieffer, 1909; 11 – A. longistilus Kieffer, 1909; 12 – A. marginalis Kieffer, 1909; 13 – A. elegantula sp. nov.; 14 – A. subtilis sp. nov. Scale = 1 mm.

Figs. 22-28. Antennae, lateral view (female). 22 – Aclista insolita Nixon, 1957; 23 – A. vernalis sp. nov.; 24 – A. dubia Kieffer, 1909; 25 – A. longistilus Kieffer, 1909; 26 – A. marginalis Kieffer, 1909; 27 – A. elegantula sp. nov.; 28 – A. subtilis sp. nov. Scale = 2 mm.

Figs. 29-42. 29-35 – fore wing venation (female). 29 – Aclista insolita Nixon, 1957; 30 – A. vernalis sp. nov.; 31 – A. dubia Kieffer, 1909; 32 – A. longistilus Kieffer, 1909; 33 – A. marginalis Kieffer, 1909; 34 – A. elegantula sp. nov.; 35 – A. subtilis sp. nov. 36-42 – proximal part of antenna, lateral view (male). 36 – A. insolita; 37 – A. vernalis sp. nov.; 38 – A. dubia; 39 – A. longistilus; 40 – A. marginalis; 41 – A. elegantula sp. nov.; 42 – A. subtilis sp. nov. Scale = 1 mm (Figs. 29-35) and 0.7 mm (Figs. 36-42).