Pamphila nubila Mabille, 1891

Zhang, Jing, Cong, Qian, Shen, Jinhui & Grishin, Nick V., 2022, Taxonomic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera), Insecta Mundi 2022 (921), pp. 1-135 : 76

Pamphila nubila Mabille, 1891 is a junior subjective synonym of Papias integra ( Mabille, 1891)

Treated by Evans as a junior subjective synonym of Cynea corisana (Plötz, 1882) (type locality Suriname), Pamphila nubila Mabille, 1891 (type locality Venezuela: Porto Cabello) is placed within Papias integra specimens in the genomic tree ( Fig. 14 ) judging from the syntype in the ZMHB collection we have sequenced. This specimen (NVG-18043C11) is a syntype, because it agrees exactly with the original description, comes from the type locality mentioned in the description according to its labels (“Pto Cabello”), has a label with “ Pa. nubila ♀ Mb. ” in handwriting similar to that of Mabille, comes from the Staudinger collection, and is curated as the type of this taxon. To stabilize nomenclature, N.V.G. hereby designates this ZMHB specimen lacking left hindwing and bearing the following labels || Origin. || Pto Cabello | Hahnel || Pa. nubila | ♀ Mb. || Pamph. | Nubila | Mab. || Coll. | Staudinger || Nubila | Mab. || GEN.PREP., | MIELKE | 1996 || [barcode image] http://coll.mfn-berlin.de/u/ | 44a060 || DNA sample ID: | NVG-18043C11 | c/o Nick V. Grishin || as the lectotype of Pamphila nubila Mabille, 1891 . Wing patterns of the lectotype, are in agreement with the genomic assessment. For instance, its paler-brown ground color (also mentioned in the original description) and pale overscaling around the veins on ventral surface of wings are like those of P.integra specimens, including the lectotype (NVG-15035E12), but differ drastically from the darker brown without paler veins pattern of C. corisana . Currently, we do not have evidence to support P.nubila as a valid taxon distinct from P.integra . For instance, in addition to the fact that the COI barcodes of the two lectotypes are 100% identical, P.nubila does not form a prominent clade in genomic trees. Therefore we propose that Pamphila nubila Mabille, 1891 is a junior subjective synonym of Papias integra ( Mabille, 1891) . With this action, being the first reviser (ICZN Code Art. 24), we also give precedence to the name integra over nubila , both names published in the same work at the same date ( Mabille 1891).

Figures

Figure 14. Genomic tree of Moncina, part 2. See Fig. 1 legend for notations.