Coleus scutellarioides (L.) Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830

Paton, Alan J., Mwanyambo, Montfort, Govaerts, Rafael H. A., Smitha, Kokkaraniyil, Suddee, Somran, Phillipson, Peter B., Wilson, Trevor C., Forster, Paul I. & Culham, Alastair, 2019, Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera, PhytoKeys 129, pp. 1-158 : 81-84

Coleus scutellarioides (L.) Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830

Ocimum scutellarioides L., Sp. Pl. ed. 2: 833. 1763.

Plectranthus scutellarioides (L.) R.Br., Prodr. Fl. Nov. Holl.: 506. 1810.

Plectranthus aromaticus Roxb., Hort. Bengal.: 45. 1814.

Majana scutellariodes (L.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.

Solenostemon scutellarioides (L.) Codd, Bothalia 11: 439. 1975.

Calchas scutellarioides (L.) P.V.Heath, Calyx 6: 51. 1999. Type: Majana (alba et rubra) Rumphius, Herb. Amboin. 5: 291, t. 101. 1747, (lectotype: illustration designated by Merrill (Interpr. Rumph. Herb. Amboin.: 460 1917).

Polypodium ovatum Burm. f., Fl. Ind.: 233. 1768. Type: Java, Unknown collector (holotype: G).

Germanea nudiflora Poir. in J.B.A.M.de Lamarck, Encycl. 2: 763. 1788.

Plectranthus nudiflorus (Poir.) Willd., Sp. Pl. 3: 168. 1800. Type: Amboina, Willdenow Herbarium 11079.1 & 11079.2 (syntypes: B-W).

Ocimum peltatum Schweigg. ex Schrank, Denkschr. Königl.-Baier. Bot. Ges. Regensburg 2: 55. 1822. Type: Cultivated from seed collected by Martius in Brazil (type: not seen).

Plectranthus ingratus Blume, Bijdr. Fl. Ned. Ind.: 836. 1826.

Coleus ingratus (Blume) Benth., Labiat. Gen. Spec.: 53. 1832.

Coleus scutellarioides var. ingratus (Blume) Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Java, G. Parang, Blume 992 (lectotype: L, designated by Keng (1969)).

Plectranthus laciniatus Blume, Bijdr. Fl. Ned. Ind.: 838. 1826.

Coleus laciniatus (Blume) Benth., Labiat. Gen. Spec.: 56. 1832.

Coleus scutellarioides var. laciniatus (Blume) Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Java, Blume s.n. (lectotype: L, designated by Keng (1969)).

Plectranthus scutellarioides Blume, Bijdr. Fl. Ned. Ind.: 837. 1826., nom. illeg.

Coleus blumei Benth., Labiat. Gen. Spec.: 56. 1832.

Coleus scutellarioides var. blumei (Benth.) Miq., Fl. Ned. Ind. 2: 950. 1858.

Majana scutellarioides var. blumei (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.

Solenostemon blumei (Benth.) M. Gómez, Bol. Secr. Agric. Comerc. Trab., Cuba 22: 127. 1914.

Plectranthus blumei (Benth.) Launert, Mitt. Bot. Staatssamml. München 7: 301. 1968. Type: Indonesia, Java, Blume s.n. (lectotype: L, designated by Suddee et al. (2004)).

Coleus atropurpureus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830.

Majana scutellarioides var. atropurpureus (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.

Calchas atropurpureus (Benth.) P.V.Heath, Calyx 5: 160. 1997. Type: Singapore, Wall. Cat. 2733A (syntypes: K (no barcode), K-W (K001116976)).

Coleus atropurpureus var. ramosus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830. Type: Malaysia, Penang, Wall. Cat. 2733B (syntypes: G-DC(microfiche), K-W (K001116976).

Coleus acuminatus Benth., Linnaea 6: 81. 1831.

Majana acuminata (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.

Calchas acuminatus (Benth.) P.V.Heath, Calyx 5: 160. 1996. Type: Philippines, Island of Samar, 1841, Cuming 1683 (neotype: K (K000897809), neotype designated by Suddee et al. 2004; isoneotypes: BM, K (K000897810), P, TCD).

Coleus atropurpureus var. densiflorus Benth., Labiat. Gen. Spec.: 54. 1832. Type: Philippines, Manila, 1830, Chamisso s.n. (syntype: K (K000897808).

Coleus grandifolius Benth., Labiat. Gen. Spec.: 53. 1832.

Majana grandifolia (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.

Coleus scutellarioides var. grandifolius (Benth.) Keng, Gard. Bull. Singapore 24: 58. 1969. Type: Timor, Guichenot s.n. (syntype P; isosyntypes: K, L, G-DC).

Coleus multiflorus Benth., Labiat. Gen. Spec.: 55. 1832.

Majana multiflora (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Philippines, Manila, Perrottet s.n. (holotype: P).

Coleus secundiflorus Benth., Labiat. Gen. Spec.: 55. 1832.

Majana secundiflora (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Timor, Guichenot s.n. (holotype: P).

Coleus grandifolius Blanco, Fl. Filip.: 482. 1837., nom. illeg.

Coleus blancoi Benth. in A.P.de Candolle, Prodr. 12: 79. 1848.

Majana blancoi (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Phillippines, Blanco, s.n. (type: not seen).

Coleus pumilus Blanco, Fl. Filip.: 482. 1837.

Majana pumila (Blanco) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Philippines, Barrio of Pineda, Pasig, Rizal Province, Luzon, Oct. 1914, Merrill: Species Blancoanae No. 190 (neotype: K, designated by Suddee et al. (2004)).

Coleus atropurpureus var. javanicus Benth. in A.P.de Candolle, Prodr. 12: 74. 1848. Type: Indonesia, Java, Zollinger 138 (syntypes: G-DC, K).

Coleus blumei var. pectinatus C.Morren, Belgique Hort. 6: 99. t 18. 1856. Type: Cultivated in Liege, illustration cited above (lectotype: illustration designated here).

Coleus scutellarioides var. celebicus Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Celebes, Reinwardt s.n. (type: not seen).

Coleus scutellarioides var. gracilis Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Ceram, Bangka, Miquel (type: not seen).

Perilla nankinensis Wender., Linnaea 29: 726. 1859, nom. illeg., non P. nankinensis (Lour.) Decne.

Coleus verschaffeltii Lem., Ill. Hort. 8: t. 293. 1861. Type: based on cultivated plants from Java,illustration cited above (lectotype: illustration, designated by Suddee et al. 2004).

Coleus marmoratus W.Bull, Proc. Roy. Hort. Soc. London 4: 133. 1864.

Coleus verschaffeltii var. marmoratus (W.Bull) André, Pl. Feuill. Ornem.: 144. 1866. Type: not seen.

Coleus gibsonii Verl., Veitch & Sons in Gard. Chron. 1866: 432 (12 May 1866). Rev. Hort. (Paris) 37: 279. 1866 (July 1866). Type: Cultivated in London, sent by John Veitch from New Caledonia. (type: not seen).

Coleus veitchii Dombrain, Fl. Mag. (London) 6: t. 345. 1867. Type: Cultivated in London, sent by John Veitch from New Caledonia, illustration cited above (lectotype: illustration, designated here).

Coleus batemannii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.

Coleus bausei T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.

Coleus berkeleyi T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.

Coleus clarkii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.

Coleus dixii T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.

Coleus marshallii T.Moore, Gard. Chron. 1868: 376. 1868.

Coleus blumei var. marshallii (T.Moore) Rothsch., Pl. Feuill. Col., 2: 74. t. 37. 1869. Type: Illustration cited above (lectotype: illustration, designated here).

Coleus murrayi T.Moore, Gard. Chron. 1868: 377. 1868.

Coleus blumei var. murrayi (T.Moore) Rothsch., Pl. Feuill. Col., 2: 74 t. 37. 1869. Type: Illustration cited above (lectotype: illustration, designated here).

Coleus reveesii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.

Coleus ruckeri T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.

Coleus saundersii T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.

Coleus scottii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.

Coleus wilsonii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.

Coleus telfordii McPhail ex H.Laurentius, Nursery Cat. (H.Laurentius) 40: 3. 1868.

Coleus blumei var. telfordii (McPhail ex H.Laurentius) Rothsch., Pl. Feuill. Col., 2: 74 t. 37. 1869. Type: Illustration cited above (lectotype: illustration, designated here).

Coleus scutellarioides var. angustifolia Benth., Fl. Austral. 5: 80. 1870.

Calchas scutellarioides var. angustifolia (Benth.) P.V.Heath, Calyx 6: 51. 1999. Type: Australia, Table Hill, Victoria River & Macadam Range, 1855, F. Mueller s.n. (syntype: K); Port Essington, Armstrong 244 (syntype: K).

Coleus scutellarioides var. laxa Benth., Fl. Austral. 5: 80. 1870. Types: Australia, Roe River, York Sound, NW coast, A.Cunningham (syntype: K); Roebuck Bay NW coast, Martin s.n. (syntype: not seen); Arnhems land, Mackinley, R.Brown (syntype: MEL).

Coleus scutellarioides var. limnophila Benth., Fl. Austral. 5: 80. 1870.

Calchas scutellarioides var. limnophila (Benth.) P.V.Heath, Calyx 6: 51. 1999. Type: Australia, Nicholson & MacArthur Rivers, F. Mueller (syntypes: K).

Coleus tryonii auct., Fl. Mag. (London), n.s., 1: t. 34. 1872. Type: cultivated in UK, Leicester, garden of Captain Tryon (lectotype: illustration designated here).

Coleus verschaffeltii var. splendens auct., Garden (London 1871-1927) 5: 54. 1874. Type: not seen.

Coleus multicolor J.H.Veitch ex Kellock, Gard. Chron., n.s., 7: 571. 1877. Type: not seen.

Coleus hendersonii Regel, Gartenflora 28: 244. 1879. Type: not seen.

Coleus x eureka auct., Gard. Chron., ser. 3, 5: 808. 1889. nom. nud.

Coleus gaudichaudii Briq., Annuaire Conserv. Jard. Bot. Genève 2: 237. 1898. Type: Philippines, fle de Luzon pres de Manille, Dec. 1836, Gaudichaud s.n. (holotype: G-DEL).

Coleus igolotorum Briq., Annuaire Conserv. Jard. Bot. Genève 2: 236. 1898. Type: Philippines, fle de Luzon, dans le pays des Igolotes, Callery 48 (holotype: G-DEL; isotype: P).

Coleus savannicola K.Schum. in K.M.Schumann & C.A.G.Lauterbach, Fl. Schutzgeb. Südsee: 529. 1900. Type: not seen.

Coleus formosanus Hayata, J. Coll. Sci. Imp. Univ. Tokyo 22: 320. 1906. Type: Kotosho, 21 Nov. 1899, Miyake s.n. (holotype: TI).

Coleus macranthus var. crispipilus Merr., Philipp. J. Sci. 1(suppl.): 235. 1906.

Coleus crispipilus (Merr.) Merr., Philipp. J. Sci., C 5: 382. 1910.

Calchas crispipilus (Merr.) P.V.Heath, Calyx 5: 160. 1996.

Coleus scutellarioides var. crispipilus (Merr.) H.Keng, Gard. Bull. Singapore 24: 56. 1969.

Calchas scutellarioides var. crispipilus (Merr.) P.V.Heath, Calyx 6: 51. 1999. Type: Philippines, Suyoc to Panai, Province of Benguet, Luzon, 2200 m, Nov. 1905, Merrill 4780 (holotype: PNH; isotypes: BM, K, NY).

Coleus pubescens Merr., Philipp. J. Sci., C 3: 432. 1909. Type: Philippines, Babuyan Island, 17 June 1907, Eugenio Zenix, Bureau of Science 3892 (holotype: PNH; isotype: BM).

Coleus zschokkei Merr., Philipp. J. Sci., C 5: 382. 1910. Type: Philippines, Mt Pulog, Prov. of Bengnet, Luzon, May 1909, Merrill 6529 (holotype: PNH; isotypes: K, NY, P).

Coleus integrifolius Elmer, Leafl. Philipp. Bot. 7: 2696. 1915.

Coleus scutellarioides var. integrifolius (Elmer) Keng, Gard. Bull. Singapore 24: 58. 1969. Type: Philippines, Mindanao, Cabadbaran (Mt Urdaneta), Prov. of Agusan, Island of Mindanao, August 1912, Elmer 13627 (holotype: PNH; isotypes: A, BM, K, L, NY, P).

Coleus gibbsiae S.Moore in L.S.Gibbs, Fl. Arfak Mts.: 178. 1917.

Coleus scutellarioides var. gibbsiae (S.Moore) Keng, Gard. Bull. Singapore 24: 58. 1969. Type: Indonesia, New Guinea, Arfak Mts, 2100 m, Gibbs 5909 (holotype: BM; isotypes: K, L).

Coleus rehneltianus A.Berger, Bot. Jahrb. Syst. 54 (5, Beibl. 120): 197. 1917. Type: not seen.

Coleus hybridus Cobeau, Arch. Bot. ( Forlì) 4: 35. 1928. Type: not seen.

Distribution.

Trop. & Subtrop. Asia to N. Australia. Widely cultivated as an ornamental.