Coptodryas Hopkins, 1915

Smith, Sarah M., Beaver, Roger A. & Cognato, Anthony I., 2020, A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China, ZooKeys 983, pp. 1-442 : 1

Coptodryas Hopkins, 1915

Coptodryas Hopkins, 1915a: 54.

Type species.

Coptodryas confusa Hopkins, 1915a; original designation.

Diagnosis.

1.8-4.0 mm, 1.88-2.71 × as long as wide. Coptodryas is distinguished by the scutellum minute, convex, slightly raised above elytral surface or not apparent; dense tuft of setae present along elytral base associated with an elytral mycangium ( C. confusa also has a pair of pit mycangia on the pronotal disc); elytral bases sinuate, costate; antennal club flattened, types 3 or 4, sutures gently sinuate and pubescent on anterior face, three sutures visible on posterior face; pronotal disc finely asperate (rarely punctate); pronotum from lateral view basic (type 0), or long and conical (type 5), rarely taller than basic (type 2; C. confusa ); pronotum from dorsal view rounded (type 1) or basic and parallel sided (type 2), rarely conical (type 0; C. confusa ); and anterior margin of pronotum with or without a row of 2-6 serrations. In addition, the procoxae are contiguous, outer margin of protibiae obliquely or distinctly triangular, armed by six or seven denticles, and posterior face flattened, unarmed.

Similar genera.

Microperus , Schedlia .

Distribution.

Species are distributed in tropical Asia and are rare in Melanesia.

Gallery system.

The gallery system in this genus appears to be rather variable ( Browne 1961b). In C. bella and C. punctipennis (Schedl, 1953), an unbranched entrance tunnel leads to a single terminal brood chamber in the longitudinal plane. In C. confusa , the tunnels are simply branched and expanded in places to form small, irregular brood chambers in the longitudinal plane. In C. quadricostata and C. curvidens (Schedl, 1958), which usually breed in small diameter stems, there is a bifurcate or circumferential gallery in the transverse plane, and one or two longitudinal branches of very variable width in which the larvae develop ( Browne 1961b).

Remarks.

Coptodryas is in need of further taxonomic/phylogenetic investigation given its potential polyphyly ( Cognato et al. 2020b) and morphological overlap with Microperus ( Hulcr et al. 2007).