Calvarium (Calvariellum) bellendenker, Zwick, Peter, 2014

Zwick, Peter, 2014, Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer, Zootaxa 3846 (1), pp. 1-41 : 12

Calvarium (Calvariellum) bellendenker , n. sp.

( Figs. 32 − 37 )

Type material. Holotype: 1♂ paratypes 3♂: “Russell R. at Bellenden Ker Landing [17.2726S, 145.9485E] 5m, 24 Oct − 9 Nov.1981 Earthwatch\ Qld.Museum” ( ANIC).

Additional material studied. 1♀ (presumed) with the same data as the types, plus “ ANIC Coleoptera Voucher No. 83-0582” ( QMSB).

Habitus. Externally indistiguishable from the congeners. BL 1.9 − 2.1 mm. Uniformly chocolate brown, appendages lighter.

Male. T8 short, strongly transverse, apodemes not continuing onto plate. Only sides of caudal margin well defined and sclerotized, middle membranous with numerous microtrichia in closely packed groups of 4 (3 − 5) ( Fig. 32 ). S8 almost unpigmented and difficult to see, approximately triangular ( Fig. 33 ).

T9 without plate, the lateral struts loosely connected by a transverse barely sclerotized crescent, apices curved mediad, with delicate apical microtrichia ( Fig. 34 ). S9 oval, with narrow, handle-like base. Colourless, distal portion finely pilose ( Fig. 35 ).

Penis with wide triangularly excised pala with sharp anterolateral corners. Pala shorter than the apical portion of penis. Trigonium with wide base, waisted, with claw-shaped centema. The flat parameroids very slender, curving around trigonium at some distance. Their widened axe-shaped tips turn mediad and meet, forming a frame around the trigonium ( Fig. 36 ).

Tegmen and parameres shaped like a slender Y, parameres basally abruptly widened and then tapering to sharply pointed tips ( Fig. 37 ).

Female (presumed). As for genus. No prehensor or vulvar sclerite found.

Note. The caudal contact between the ends of the parameroids distantly resembles the structure of Austrocyphon spp. (Zwick 2013c). The female is very probably conspecific with the males with which it was taken. However, it cannot unambiguously be identified. The female is therefore assigned no type status.

Etymology. Named after the type locality over which Mt Bellenden Ker peaks, the second highest mountain in Queensland (1557m).

Figures

FIGURES 29 − 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale.