Calvarium (Calvariellum) hamifer, Zwick, Peter, 2014

Zwick, Peter, 2014, Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer, Zootaxa 3846 (1), pp. 1-41 : 9-10

Calvarium (Calvariellum) hamifer , n. sp.

( Figs. 6 − 9 , 18 − 27 )

Type material. Holotype ♂: “ 12.44S 143.148E QLD 3km ENE Mt. Tozer 28 June − 4 July 1986 D.H.Colless \ Malaise trap” ( ANIC). Paratypes: 2♂ with the same data; 2♂ with the same data, except 11 − 16 July 1986; 1♂ “ 15.30S 145.16E 1km SE of Mt. Cook QLD 12.Oct.1980 D.H.Colless (at light)”; 2♂ “ 15.30S 145.16E Mt. Cook Ntl Pk QLD 13.Oct.1980 D.H.Colless”; 2♀: “ 15.19S 145.03E Station Ck 7 km WSW Hope Vale Mission QLD 10.May 1981 A.Calder \ by sweeping”; 4♂ 5♀: “Rocky Ck., 7 M. N of Atherton N.Qld. 3 May 1967 D.H.Colless“; 4♂ 1♀: “Bamboo Ck., near Miallo N. of Mossman N.Qld. 25 Apr. 1967 D.H. Colless“; 2♂ 1♀: “ 7 − 14 m. W. of Herberton, via Watsonville N.Qld. 1 May 1967 D.H.Colless“; 1♀: “Millstream at Archer's Ck, N.Q., Mt. Garnet Rd. 28.xii.64, J.G.Brooks“; 1♂: “ 16.55S 145.52E Yarrabah, Q. 17.v.76, E.B.Britton“ (all ANIC). 1♂: “Upper Cedar Ck., via Samford [27° 22.333' S 152° 53.184' E], QLD. 6.xii.62 G. Monteith” \ UQIC Reg. #88682 ( QMSB); 2♂ 1♀: “Sth. Pine R., 17.i.63 QLD. G. Monteith” \ UQIC Reg. #88665, #88669, #88670 ( QMSB). 1 ♀, “Ellis Beach 12/10/12 C.Watts“ ( SAMA).

Habitus. BL 1.9 − 2.2 mm, BL /BW ~ 1.7. Dark brown, legs lighter brown, antennae yellowish, distal half infuscate.

Male. S7 a little pointed. T8 narrow, caudal edge slightly angular. Only grouped microtrichia on disc, no setae. To the inside of each apodeme arises on the ventral face a long sclerotized sharply tipped blade ( Fig. 21 ). S8 is an unpigmented hairless caudally excised small plate with short paired apodemes ( Fig. 22 ). T9 represented by two slender slightly sinuous long rods with sharp caudal tip. S9 unpigmented, base slender, unpaired, the oval plate caudally pilose ( Fig. 23 ).

Pala a wide ventrally open tubular trough, its base broadly excised between prominent anterolateral corners. Pala narrowing caudad, a little longer than the narrower distal part of the penis. Trigonium plump, finger-shaped with slender basal spur and caudally with a huge downcurved hollow hook. Parameroids shorter than trigonium, slender, widening and slightly curved outward in distal half, apices rounded with small sensory pores ( Fig. 24 ). Tegmen a small U-shaped sclerite supporting the parameres which are first abruptly widened and then continuously narrowed to a sharp tip ( Fig. 25 ).

Female. Ovipositor as for genus ( Fig. 26 ). Prehensor complex ( Fig. 27 ), with a deep sleeve-like circular fold and unarmed bare folded sclerites.

Note. The large blades on the lower face of T8 are exceptional. When during dissection of macerated specimens the terminal segments were pulled out T8 was rising until its disc folded forward while the blades on the lower face of T8 pointed back. C. (Calvariellum) cochlearifer n. sp. possesses in the same place small outgrowths that appear like forerunners of the present condition. The species is known from far NE Queensland to South Pine River NW of Brisbane. No congeners are known so far South which helped association of sexes.

Etymology. The name is a noun in apposition, formed from Latin hamus, hook and ferre, to carry: the hookcarrier.

Figures

FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture.

FIGURES 18 − 20. Calvarium (Calvariellum) hamifer, n. sp., male: 18, contour in dorsal view; 19, antenna; 20, central portion of thorax, ventral side. After photographs. Fig. 20 not to scale, diagrammatic. Abbreviations: cx I, fore coxa; cx II, middle coxa; g, groove receiving prosternal process (psp); msp, process of mesoventrite; mtp, process of metaventrite.

FIGURES 21 − 25. Calvarium (Calvariellum) hamifer, n. sp., male: 21, T 8, ventral; 22, S 8; 23, S 9 and the apodemes representing T 9, superimposed; 24, penis; 25 tegmen (25 holotype, the other paratypes from Mt. Tozer; all to the same scale).

FIGURES 26 − 28. Calvarium (Calvariellum) spp., females: 26, C. hamifer, n. sp. (paratype from Station Ck), abdomen with protracted ovipositor and prehensor in situ; 27, prehensor enlarged, diagrammatic. 28, C. cochlearifer, n. sp. (from Bamboo Ck), prehensor. Abbreviation: cf, circular fold. 27 and 28 to the same scale.