Ypsiloncyphon

Zwick, Peter, 2014, Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer, Zootaxa 3846 (1), pp. 1-41 : 19-20

Key to females of species group 3 of Ypsiloncyphon

(females of Y. pusillus , Y. rugosus , Y. velatus , and Y. virgulifer unknown)

1 Apodemes of S8 anteriorly fused, forming a single median sclerite (e.g., Figs. 68, 88, 106)........................... 2

1' Apodemes of S8 connected in front to form an open frame of various forms (e.g., Figs. 116 , 119 , 124 ).................. 7

2 Unpaired apodeme of S8 wide, a slender triangular sclerite caudally drawn out into the lateral rods, anteriorly with a middorsal tooth ( Fig. 127 ).................................................................................. Y. sp. A

2' Sclerites of S8 resembling a tuning fork, the unpaired apodeme is a slender rod without tooth......................... 3

3 T7 with two paramedian notches ( Fig. 107 ). Vulvar sclerite box-shaped, dictyon composed of few large meshes ( Figs. 108, 109 )................................................................................ Y. katherinae , n. sp.

3' T7 without notches. S7 with median nipple which is folded forward, not visible from outside. Vulvar sclerite widening posteri- orly, a dorsally widely open tube ending in two spreading caudolateral lobes (e.g., Figs. 82, 89); dictyon large, composed of many meshes (e.g., Fig. 83)............................................................................. 4

4 Unpaired portion of apodemes of S8 short, less than half as long as the paired branches ( Fig. 129 ). Conspicuous scales cau- dolaterally on vulvar sclerite ( Fig. 131 )............................................................... Y. sp. B

4' Unpaired and paired parts of apodemes of S8 of similar length. Vulvar sclerite not scaly............................. 5

5 Vulvar sclerite weak, two elongate parallel sclerites caudally connected by a short transverse sclerite bridge (Fig. 68)............................................................................................. Y. ruficollis , n. sp.

5' Vulvar sclerite large, resembling a box with dorsal slit........................................................ 6

6 Paramedian crests of vulvar sclerite close together, caudal lobes hard, sharply pointed (Fig. 82)............ Y. longus , n. sp.

6' Paramedian crests of vulvar sclerite more widely separate, caudal lobes soft, lappet-like ( Fig. 89 )........ Y. mutilatus , n. sp.

7 Anterior ends of apodemes of S8 form a regular oval frame (e.g., Figs. 116 , 132 )................................... 8

7' Apodemes of S8 form an anteriorly widening frame with large sclerotized expansions along edge ( Figs. 119 , 124 )......... 9

8 Vulvar sclerite a soft elongate body with several lobes on both ends ( Fig. 133 )................................ Y. sp. C

8' Vulvar sclerite with transverse sclerotized bar with short forward extensions on the sides ( Fig. 116 ).. Y. micans (Klausnitzer)

9 Apodemes of S8 trapezoid with sclerotized expansions forming anterior wings extending forward beyond the delicate trans- verse connection between them, vulvar sclerite tube 5–6 times longer than wide ( Fig. 124 ). T7 with small mediocaudal out- growth ( Fig. 125 )........................................................................ Y. angustus , n. sp.

9' Lateral expansions to apodemes of S8 distinctly behind oval front loop, vulvar sclerite tube at most 4 times longer than wide ( Fig. 119 ). T7 unmodified.................................................................... Y. brevis , n. sp.

Figures

FIGURES 110 – 116. Ypsiloncyphon micans Klausnitzer. 110, 111 penis tip and S 9 of the Y. micans holotype. 112 – 115, holotype of C. paramicans: 112, T 8; 113, T 9; 114, penis; 115, S 9; 116, presumed female, S 8 and vulvar sclerite. Figs. 112 – 116 to the same scale.

FIGURES 117 – 120. Ypsiloncyphon brevis, n. sp.: 117, male S 9; 118, penis, dorsal (Sth. Pine R.); 119, female terminal segments with vulvar sclerite, apodemes of S 8 black, expansions grey (Goldmine & Davies Cks); 120, dictyon. All to the same to scale.

FIGURES 121 – 126. Ypsiloncyphon angustus, n. sp.: 121, male S 9; 122, penis, dorsolateral (near Herberton); 123, penis, dorsal (Big Mitchell Ck); 124, female terminal segments with vulvar sclerite, apodemes of S 8 black, expansions grey (Rocky Ck); 125, female, tip of T 7; 126, dictyon. Figs. 121 – 123 to the same scale, 125 not to scale.

FIGURES 127, 128. Ypsiloncyphon sp. A: 127, part of female terminalia, apodemes of S 8 (black mark points at the triangular process), vulvar sclerite in center; 128, vulvar sclerite, detail. Not to scale.

FIGURES 103 – 109. Ypsiloncyphon katherinae, n. sp., male (103 – 105) and female (106 – 109): 103, ♂ holotype, S 9; 104, penis, dorsal; 105, tegmen and parameres; 106, ♀ (N of Tennant Ck.) terminal segments, apodemes of S 8 black; 107, ♀ T 7; 108, vulvar sclerite; 109, dictyon. Figs. 103 – 106 to the same scale.

FIGURES 129 – 133. Ypsiloncyphon sp. B (129 – 131; from near King Cascade) and Y. sp. C (132, 133); 129, female terminal segments, with vulvar sclerite in situ; apodemes of S 8 black; 130, dictyon; 131, vulvar sclerite, enlarged, not to scale. 132, female segment 8, apodemes of S 8 black, flange grey; 133, vulvar sclerite. 129 & 130, and 132 & 133 to the same scale, respectively.

FIGURES 84 – 89. Ypsiloncyphon mutilatus, n. sp. (paratypes). 84, male T 8 and T 9, superimposed; 85, tegmen and parameres; 86, penis in oblique view. 87, penis, dorsal view; 88, female terminal segments with vulvar sclerite; 89, vulvar sclerite. 84 – 87 to the same scale.