Ypsiloncyphon

Zwick, Peter, 2014, Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer, Zootaxa 3846 (1), pp. 1-41 : 19

Key to males of species group 3 of Ypsiloncyphon

1 Parameroids with curved subterminal appendage ( Fig. 60 )....................................... Y. virgulifer , n. sp.

1' Parameroids without appendage.......................................................................... 2

2 Head and pronotum bright red, rest of body dark brown......................................... Y. ruficollis , n. sp.

2' Dorsal side dark brown to black, at most head reddish......................................................... 3

3 Outer face of parameroids covered with small hard teeth, rough ( Fig. 73 ). S9 composed of a pair of similar, thin, pointed scler- ite rods ( Fig. 72 )......................................................................... Y. rugosus , n. sp.

3' Parameroids with smooth surface. S9 pronouncedly asymmetrical, caudal parts blade-like............................ 4

4 Pala several times longer than the short capulus of S9. S9 about 50% longer than the penis (Figs. 77, 79).... Y. longus , n. sp.

4' Capulus and pala of similar length (e.g., Figs. 64, 66, 103, 104). Penis never so much shorter than S9................... 5

5 Parameroids pointed, ending in a thin flagellum. Parameroids about 1/3 longer than the trigonium ( Fig. 86 ) Y. mutilatus , n. sp.

5' Parameroids a little (ca. 10–15%) longer than the trigonium, their apices blunt..................................... 6

6 Pala long, forming a long rod-like base to the variable trigonium (e.g., Fig. 93 , 101 )................................. 7

6' Pala waisted, constricted portion short, not a long narrow stalk ( Figs 118 , 123 ). Trigonium not waisted, gradually narrowing to tip................................................................................................. 10

7 Short blade of S9 apically abruptly narrowed, narrow portion a little longer than maximum width of blade (e.g., Figs 94 , 102 ).................................................................................................... 8

7' Short blade of S9 gradually drawn out into a narrow apical portion which is two or more times longer than maximum width of blade (e.g., Figs. 103 , 115 ).............................................................................. 9

8 Parameroids apically wide, enclosing the trigonium between them, veiling it partly. Trigonium waisted, narrowest point of similar width as pala at its narrowest point ( Figs. 93, 95, 97 )....................................... Y. velatus , n. sp.

8' Parameroids slender, trigonium exposed, its narrowest point only about half as wide as narrowest point of pala ( Fig. 101 ).......................................................................................... Y. pusillus , n. sp.

9 Trigonium distinctly waisted, the apex wide, bluntly rounded, with small median nipple. Parameroids are sinuous rods. Narrow apical portion of both blades of S9 curved ( Figs. 103, 104 )...................................... Y. katherinae , n. sp.

9' Trigonium conical, continuously narrowed to tip, not constricted ( Fig. 114 ). Parameroids are slender plates. Blades of S9 drawn out into long straight apices ( Figs. 111, 115 )......................................... Y. micans (Klausnitzer)

10 Penis short, stout, straight, trigonium about 4–5 times longer than maximum width, its tip widely separate from the param- eroids ( Fig. 118 )........................................................................... Y. brevis , n. sp.

10' Penis long and slender, trigonium up to 7 times longer than maximum width, trigonium and parameroid tips close together, trigonium tip often bent back ( Figs. 122, 123 )................................................. Y. angustus , n. sp.

Figures

FIGURES 58 – 61. Ypsiloncyphon virgulifer, n. sp., male: 58, T 8, T 9 and S 9, superimposed; 59, penis, ventral view; 60, oblique dorsolateral view of same, trigonium tip bent ventrad; 61, tegmen and parameres. Pilosity and surface structure of T 8 and T 9 shown only diagrammatically in small selected areas. All to same scale.

FIGURES 70 – 74. Ypsiloncyphon rugosus, n. sp., male holotype: 70, T 8; 71, T 9; 72, S 9; 73, penis; 74, tegmen and parameres. All to same scale.

FIGURES 84 – 89. Ypsiloncyphon mutilatus, n. sp. (paratypes). 84, male T 8 and T 9, superimposed; 85, tegmen and parameres; 86, penis in oblique view. 87, penis, dorsal view; 88, female terminal segments with vulvar sclerite; 89, vulvar sclerite. 84 – 87 to the same scale.

FIGURES 90 – 98. Morphs of Ypsiloncyphon velatus, n. sp.: 90, T 8; 91, T 9; 92, tegmen and parameres; 93, 95, 97, penis; 94, 96, 98, S 9. All to same scale. 93, 94, Morph 2 from Cadjeput Rockhole; 95, 96, holotype; 97, 98, Morph 1 from near Oenpelli.

FIGURES 99 – 102. Ypsiloncyphon pusillus, n. sp., male holotype: 99, T 8; 100, T 9; 101, penis lying over tegmen and parameres; 102, S 9. 99 & 100, and 101 & 102 to the same scales, respectively.

FIGURES 117 – 120. Ypsiloncyphon brevis, n. sp.: 117, male S 9; 118, penis, dorsal (Sth. Pine R.); 119, female terminal segments with vulvar sclerite, apodemes of S 8 black, expansions grey (Goldmine & Davies Cks); 120, dictyon. All to the same to scale.

FIGURES 121 – 126. Ypsiloncyphon angustus, n. sp.: 121, male S 9; 122, penis, dorsolateral (near Herberton); 123, penis, dorsal (Big Mitchell Ck); 124, female terminal segments with vulvar sclerite, apodemes of S 8 black, expansions grey (Rocky Ck); 125, female, tip of T 7; 126, dictyon. Figs. 121 – 123 to the same scale, 125 not to scale.

FIGURES 103 – 109. Ypsiloncyphon katherinae, n. sp., male (103 – 105) and female (106 – 109): 103, ♂ holotype, S 9; 104, penis, dorsal; 105, tegmen and parameres; 106, ♀ (N of Tennant Ck.) terminal segments, apodemes of S 8 black; 107, ♀ T 7; 108, vulvar sclerite; 109, dictyon. Figs. 103 – 106 to the same scale.

FIGURES 110 – 116. Ypsiloncyphon micans Klausnitzer. 110, 111 penis tip and S 9 of the Y. micans holotype. 112 – 115, holotype of C. paramicans: 112, T 8; 113, T 9; 114, penis; 115, S 9; 116, presumed female, S 8 and vulvar sclerite. Figs. 112 – 116 to the same scale.