Czernyola, Bezzi

Owen Lonsdale, 2017, World Catalogue of the Druid Flies (Diptera: Schizophora: Clusiidae), Zootaxa 4333 (1), pp. 1-85 : 29-35

publication ID	https://doi.org/ 10.11646/zootaxa.4333.1.1
publication LSID	lsid:zoobank.org:pub:47AB9D4D-16FF-4A7E-B73F-2A65C2841FF4
DOI	https://doi.org/10.5281/zenodo.4752271
persistent identifier	https://treatment.plazi.org/id/103F3650-BF67-FFB0-FF0C-FA13DA40FCBA
treatment provided by	Plazi
scientific name	Czernyola
status

Treatment

Genus CZERNYOLA Bezzi View in CoL

Craspedochaeta Czerny 1903: 103 View in CoL [preoccupied by Macquart (1851), which was originally spelled Craspedochoeta , but corrected to Craspedochaeta by Scudder (1882)]. Type species, Craspedochaeta transversa Czerny 1903 , subsequent designation by Melander & Argo (1924: 24). Gender feminine. Kertész 1903: 569 [review, key], 1906: 320 [key]; Schulze et al. 1928: 818 [nomenclator]; Neave 1939a: 860 [nomenclator]; McAlpine 1971: 121 [synonymy of Czernyola Bezzi View in CoL and Chaetoczernyola Soós ]; Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific]; Sueyoshi 2006: 12 [ Japan], 2014: 618 [catalogue, Japan]; Papp et al. 2006: 190 [unidentified and tentatively identified species, Thailand]; Lonsdale & Marshall 2006b: 9 View Cited Treatment [New world revision, phylogeny, biology, male agonistic behaviour], 2007b: 119 [use in analysis of Clusiodes View in CoL ], 2008b: 5 [ Fiji], 2010: 1044 [Manual of Central American Diptera View in CoL ]; Lonsdale 2009b: 230 [key to genera in Australia]; Sasakawa 2010: 13 [Oriental and Pacific revision]; Evenhuis et al. 2016: 49 [as synonym of Czernyola Bezzi View in CoL ].

Czernyola Bezzi 1907: 52 View in CoL [replacement name for Craspedochaeta ]. Type species, Craspedochaeta transversa Czerny 1903 , subsequent designation by Melander & Argo (1924: 24). Gender feminine. Melander & Argo 1924: 24 [family and genus revision, type designation]; Schulze et al. 1928: 912 [nomenclator]; Curran 1934b: 364 [key, North America]; Hennig 1938: 130 [review, world], 1952: 227 [discussion of immature stages in Clusiidae View in CoL ]; Neave 1939a: 957 [nomenclator]; Malloch 1942: 209 [synonymy of Tonnoiria Malloch , Guam]; Frey 1960: 21 [family and genus revision, key, Indoaustralia, catalogue, world]; McAlpine 1960: 76 [revision, key to species and species groups, Australia]; 1971: 121 [as synonym of Craspedochaeta ]; Soós 1962c: 449 [genus revision], 1964a: 6 [key], 1968: 3 [catalogue, Americas south of USA]; Cole & Schlinger 1969: 426 [key to genera of Clusiidae View in CoL ]; Sasakawa 1971: 50 [revision, Orient/Pacific], 1977: 237 [catalogue, Orient]; Lonsdale et al. 2010: 267 [stat. reinst., family revision, key]; Lonsdale 2014: 212 [ Sobarocephala View in CoL sp. from Guam reported in Bohart & Gressit (1951) identified as Czernyola View in CoL sp.]; Evenhuis et al. 2016: 49 [as n. stat.].

Tonnoiria Malloch 1929a: 98 View in CoL . Type species, Tonnoiria palliseta Malloch 1929 , by original designation. Gender feminine. Malloch 1932: 215 [Marquesas Isl., notes undescribed species from Guam]; Neave 1940b: 508 [nomenclator]; Malloch 1942: 209 [as synonym of Czernyola View in CoL ]; Hennig 1952: 227 [discussion of immature stages in Clusiidae View in CoL ]; Steyskal & Sasakawa 1966: 244 [key, Micronesia]; Steyskal 1968: 87 [priority of Tonnoiria Malloch over Tonnoiria Parent (Dolichopodidae) ].

Praspedochaeta. Misspelling of Craspedochaeta . Frey 1960: 26.

Chaetoczernyola Soós 1962a: 141 . Type species, Chaetoczernyola novaeguineae Soós 1962 , by original designation. Gender feminine. Soós 1964a: 6 [key]; Sasakawa 1971: 62 [revision, Orient/Pacific]; McAlpine 1971: 121 [as synonym of Craspedochaeta ].

amazonensis (Lonsdale & Marshall) View in CoL . NT: Brazil (Amazonas). NEW COMBINATION

Craspedochaeta amazonensis Lonsdale & Marshall 2006b: 30 View in CoL View Cited Treatment . Brazil. Amazonas: 26km NE Manaus, Reserva Ducke. HT ♂ INPA. Lonsdale & Marshall 2006b: 30 View Cited Treatment [illustrations of male genitalia, phylogeny].

annulipes (Johnson) View in CoL . NE: United States (Maryland, Mississippi, North Carolina, Texas). NT: Tropical Neotropics. Heteromeringia annulipes Johnson 1913b: 99 View in CoL . USA. North Carolina: Hertford Co., Murfreesboro. HT ♀ MCZ. Malloch 1918: 8 [synonymy, type data]; Melander & Argo 1924: 29 [summary of original description]; Brimley 1938: 378 [North Carolina]; Frey 1960: 27 [catalogue, world]; Steyskal 1965: 806 [catalogue, America north of Mexico]; Poole 1996: 126 [checklist, Nearctic].

Czernyola basalis brasiliensis Soós 1961: 399 . Brazil. Mato Grosso: Maracaju. HT ♀ USNM. Soós 1962c: 449 [key, review, world], 1968: 3 [catalogue, Americas south of USA];; Lonsdale & Marsjall 2006b: 59 [as synonym of annulipes Johnson View in CoL ].

Czernyola piceoflava Sóos 1962a: 146 View in CoL . Dominica. Trujillo: Colonia Ranfis. HT ♂ USNM. Soós 1962c: 449 [key, review, world], 1968: 4 [catalogue, Americas south of USA]. Steyskal 1973: 443 [checklist, Antilles]; Lonsdale & Marshall 2006b: 59 [as synonym of annulipes Johnson View in CoL ].

Craspedochaeta annulipes . Lonsdale & Marshall 2006b: 59 [redescription, generic combination, synonymy of piceoflava Sóos View in CoL and basalis brasiliensis Sóos , phylogeny, new records ( Bolivia, Brazil, British Guiana, Costa Rica, Dominican Republic, Ecuador, Guyana, Honduras, Jamaica, Mexico, Panama, Peru, St. Vincent, United States (Maryland, Mississippi, Texas), U.S. Virgin Isl., Venezuela), illustrations of female and male genitalia]. Czernyola annulipes View in CoL . Lonsdale et al. 2010: 240 [generic combination, genes sequenced for phylogenetic analysis]

apsilutea (Lonsdale & Marshall) View in CoL . NT: Bolivia, Dominican Republic, Ecuador. NEW COMBINATION Craspedochaeta apsilutea Lonsdale & Marshall 2006b: 19 . Dominican Republic. Pedernales: 26km N Cabo Rojo, 565m. HT ♂ DEBU. Lonsdale & Marshall 2006b: 19 [phylogeny, illustrations of colouration and male and female genitalia].

argoniae (Lonsdale & Marshall) View in CoL . NT: Bolivia, Ecuador. NEW COMBINATION

Craspedochaeta argoniae Lonsdale & Marshall 2006b: 75 View in CoL View Cited Treatment . Ecuador. Napo Province: 7km S Baeza, 2000m. HT ♂ USNM. Lonsdale & Marshall 2006b: 75 View Cited Treatment [phylogeny, illustrations of head and male genitalia].

atrifrons Malloch. AU: Mariana Isl.

Czernyola atrifrons Malloch 1942: 209 View in CoL . Guam. Dededo. HT ♀ BPBM [Type No. 2327]. Frey 1960: 22 [redescription, catalogue, world]; Soós 1962c: 449 [key, review, world]; Sasakawa: 1971: 52 [synonymy, diagnosis]. Tonnoiria atrifrons . Steyskal & Sasakawa 1966: 245 [generic combination, illustrations of head and thorax]. Craspedochaeta atrifrons . Pitkin & Evenhuis 1989: 535 [generic combination, catalogue, Australasia/Pacific].

australis McAlpine. AU: Australia (New South Wales).

Czernyola australis McAlpine 1960: 78 View in CoL . Australia. New South Wales: Deep Creek, Narrabeen, near Sydney. HT ♂ USNM. McAlpine 1960: 78 [illustration of antenna]; Soós 1962c: 449 [key, review, world]. Craspedochaeta australis . Pitkin & Evenhuis 1989: 535 [generic combination, catalogue, Australasia/Pacific].

basalis Czerny. View in CoL NT: Costa Rica, Peru.

Craspedochaeta basalis Czerny 1903: 104 View in CoL . “Peru”. ST ♂ ♀ NMW [1♀ found in NMW; location of remaining ST, at least including 1♂, is unknown]. Kertész 1903: 569 [redescription, Peru] ; Lonsdale & Marshall 2006b: 64 View Cited Treatment [redescription, phylogeny, Costa Rica, illustration of female genitalia].

Czernyola basalis View in CoL . Melander & Argo 1924: 26 [generic combination, summary of original description]; Frey 1960: 26 [catalogue, world].

Czernyola basalis basalis View in CoL . Soós 1961: 401 [new status], 1962c: 449 [key, review, world], 1968: 3 [catalogue, Americas south of USA].

biloba (Lonsdale & Marshall) . NT: Brazil ( Amazonensis ). NEW COMBINATION

Craspedochaeta biloba Lonsdale & Marshall 2006b: 67 View in CoL View Cited Treatment . Brazil. Amazonas: Rio Préto, a tributary of Rio Pasanuari (left tributary of Rio Negro) near Padanaque, rainforest, 00°9.47′S, 67°7.19′W. HT ♂ ZSBS. Lonsdale & Marshall 2006b: 67 View Cited Treatment [phylogeny, illustration of male genitalia].

biseta (Hendel) . AU: Bismarck Isl., Papua New Guinea. OR: Japan (Ryukyus), Laos, Malaysia ( Sarawak), Philippines, Taiwan, Thailand.

Czernyola biseta Hendel 1913: 80 View in CoL . Taiwan. Kosempo and Kankau. ST 2♂ NMW [only 1♂ found in collection]. Hennig 1938: 130 [redescription]; Melander & Argo 1924: 26 [summary of original description]; Hennig 1941: 146 [ Taiwan]; Frey 1960: 22 [summary of original description, catalogue, world]; Soós 1962c: 449 [key, review, World]; Sasakawa 1971: 53 [redescription, synonymy of puncticornis Frey View in CoL and puncticornis laetior Frey View in CoL , Philippines, New Guinea]; Balthazar 1990: 417 [catalogue, Philippines, incorrect type depository listed]; Sasakawa & Pong 1990: 57 [Sabah (also recorded from Sarawak, Taiwan, Philippines, New Britain, New Guinea and New Ireland)]; Sueyoshi 2012: 26 [ Japan].

Czernyola puncticornis Frey 1928b: 106 View in CoL . Philippines. Luzon: Los Banos. HT ♀ UZMH. Frey 1928b: 106 [with catalogue]; Delfinado & Hardy 1971: 29 [catalogue Philippine types]; Sasakawa 1971: 53 [as synonym of biseta Hendel View in CoL ]; Balthazar 1990: 417 [catalogue, Philippines].

Czernyola puncticornis laetior Frey 1960: 22 View in CoL . Taiwan. HT ♀, location unknown [originally deposited in UZMH]. Frey 1960: 22 [catalogue, world]; Sasakawa 1971: 53 [as synonym of biseta Hendel View in CoL ], 1977: 237 [as synonym of biseta Hendel View in CoL , catalogue, Orient]..

Craspedochaeta biseta . Sasakawa 1974: 161 [generic combination, Philippines, Bismarck Isl.], 1977: 237 [catalogue, Orient]; Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific]; Sueyoshi et al. 2009: 100 [log emergence trap, tentative identification, Japan]; Sasakawa 2010: 15 [Oriental/Pacific revision, Laos, Thailand]; Sueyoshi 2011: 9 [ Japan, pictoral key], 2014: 618 [catalogue, Japan].

Czernyola bisela View in CoL . Misspelling of biseta . Hua 2006: 160 [catalogue—not known from China].

bisignata McAlpine. AU: Australia (Queensland). OR: Malaysia (Sarawak).

Czernyola bisignata McAlpine 1960: 78 View in CoL . Australia. North Queensland: Kuranda. HT ♂ AMS. Soós 1962c: 449 [key, review, world]; Sasakawa 1971: 54 [redescription, Sarawak], 1977: 237 [catalogue, Orient]. Craspedochaeta bisignata . Pitkin & Evenhuis 1989: 535 [generic combination, catalogue, Australasia/Pacific].

boettcheri Frey. OR: Malaysia (Samar), Philippines.

Czernyola Boettcheri Frey 1928b: 105 View in CoL . Malaysia. Samar: Catbalogan. HT ♂ UZMH.

Czernyola boettcheri View in CoL . Frey 1960: 22 [summary of original description, illustration of head, catalogue, world, Philippines]; Soós 1962c: 449 [key, review, world]; Delfinado & Hardy 1971: 29 [catalogue Philippine types, type erroneously listed as ♀]; Sasakawa 1971: 54 [synonymy, diagnosis], 1977: 238 [catalogue, Orient]; Balthazar 1990: 417 [catalogue, Philippines].

Craspedochaeta boettcheri . Lonsdale & Marshall 2006b: 5 [generic combination, phylogenetic analysis, Philippines].

borneoensis Sasakawa. OR: Malaysia (Pen., Sabah), Vietnam.

Czernyola borneoensis Sasakawa 1971: 54 View in CoL . Malaysia. Sabah: Paring, nr Ranau. HT ♂ BPBM. Sasakawa 1971: 54 [illustrations of male genitalia], 1977: 238 [catalogue, Orient]; Sasakawa & Pong 1990: 58 [illustration of male genitalia, Sabah].

Craspedochaeta borneoensis . Sasakawa 2010: 16 [Oriental/Pacific revision, Malaysia, Vietnam].

brunneivibrissa (Lonsdale & Marshall) . NT: Jamaica. NEW COMBINATION

Craspedochaeta brunneivibrissa Lonsdale & Marshall 2006b: 66 View in CoL View Cited Treatment . Jamaica. Trel. Parish ca 6 km N Quick Step near Come Night Cave. HT ♂ USNM. Lonsdale & Marshall 2006b: 66 View Cited Treatment [phylogeny, illustration of male genitalia].

caligula Sasakawa. AU: Papua New Guinea. OR: Malaysia [Pen.].

Czernyola caligula Sasakawa 1971: 55 View in CoL . Papua New Guinea. Morobe Province: Wampit V., nr Wau, Gurakor Village, 950 m. HT ♂ BPBM. Sasakawa 1971: 55 [illustrations of male genitalia].

Craspedochaeta caligula . Sasakawa 1974: 161 [generic combination, Bismarck Isl.]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific]; Sasakawa 2010: 16 [Oriental/Pacific revision, Malaysia].

candida (Lonsdale & Marshall) View in CoL . NT: Bolivia, Brazil (Roraima). NEW COMBINATION

Craspedochaeta candida Lonsdale & Marshall 2006b: 34 View in CoL View Cited Treatment . Bolivia. La Paz Department: San Silverio, 20 km S Caranvi, 15°50′S 67°36′W, 750m. HT ♂ ANCB. Lonsdale & Marshall 2006b: 34 View Cited Treatment [phylogeny, illustrations of colouration and male genitalia].

chauliodon (Lonsdale & Marshall) View in CoL . NT: Ecuador. NEW COMBINATION

Craspedochaeta chauliodon Lonsdale & Marshall 2006b: 69 View in CoL View Cited Treatment . Ecuador. Napo Province: Yasuni National Park, PUCE Yasuni Research Station, 00°38′S, 76°36′ W. HT ♂ ZSBS. Lonsdale & Marshall 2006b: 69 View Cited Treatment [phylogeny, illustrations of male genitalia].

chela (Lonsdale & Marshall) View in CoL . NT: Belize *, Costa Rica, Ecuador, Guatemala, Peru. NEW COMBINATION Craspedochaeta chela Lonsdale & Marshall 2006b: 36 . Costa Rica. Limón Province: Valle de la Estrella, R.R. Hitoy Cerere, 100– 200 m. HT ♂ INBio. Lonsdale & Marshall 2006b: 36 [phylogeny, illustrations of colouration and male genitalia].

claripennis Melander & Argo. NT: Costa Rica, Ecuador.

Czernyola claripennis Melander & Argo 1924: 26 View in CoL . Costa Rica. Cartago Province: La Suiza de Turrialba. HT ♂ USNM. Frey 1960: 27 [catalogue, world]; Soós 1962c: 449 [key, review, world], 1968: 3 [catalogue, Americas south of USA]. Craspedochaeta claripennis . Lonsdale & Marshall 2006b: 31 [generic combination, redescription, phylogeny, new records ( Costa Rica, Ecuador), illustration of female genitalia].

concinna (Williston) View in CoL . NE: United States (Florida, New Mexico). NT: Neotropics. NEW COMBINATION Heteroneura concinna Williston 1896: 387 . St. Vincent. “Leeward side”. LT ♂ BMNH [designated by Lonsdale & Marshall 2006b: 52]. Czerny 1903: 101 [quotes description, notes]; Aldrich 1905: 570 [catalogue, North America]. Craspedochaeta atra Kertész 1903: 570 . Bolivia. “Songo” [= La Paz Department: Zongo]. HT ♀ SMT. Lonsdale & Marshall 2006b: 52 [as synonym of concinna Williston ].

Czernyola atra . Melander & Argo 1924: 25 [redescription, generic combination, photo, Costa Rica]; Hennig 1938: 131 [type information]; Frey 1960: 26 [catalogue, world]; Soós 1962c: 449 [key, review, world], 1968: 3 [catalogue, Americas south of USA].

Sobarocephala concinna . Melander & Argo 1924: 36 [quotes original description, generic combination]; Frey 1960: 27 [catalogue, world]; Soós 1968: 5 [catalogue, Americas south of USA]; Steyskal 1973: 443 [checklist, Antilles]. Czernyola minuta Soós 1962a: 148 . Costa Rica. Higuto, San Mateo. HT ♂ USNM. Soós 1962c: 449 [key, review, world], 1968: 4 [catalogue, Americas south of USA]; Lonsdale & Marshall 2006b: 52 [as synonym of concinna Williston ]. Czernyola albohalteria Soós 1962a: 145 . Bolivia. Cachuela Esperanza. HT ♂ USNM. Soós 1962c: 449 [key, review, world], 1968: 3 [catalogue, Americas south of USA]; Lonsdale & Marshall 2006b: 52 [as synonym of concinna Williston ].

Craspedochaeta concinna . Lonsdale & Marshall 2006b: 52 [redescription, synonymy of atra Kertész , minuta Soós and albohalteria Soós , generic combination, discussion of pale and dark “phases”, phylogeny, new records ( Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Grenada, Guatemala, Guyana, Honduras, Jamaica, Mexico, Panama, Peru, St. Vincent, Trinidad, Venezuela, United States (Florida, New Mexico)), illustrations of head and male genitalia, photo of male and female in copula], 2010: 1046 [reproduction of illustration of female genitalia].

delta McAlpine. AU: Australia (Queensland), New Guinea.

Czernyola delta McAlpine 1960: 79 View in CoL . Australia. North Queensland: Lake Placid (or Barron Waters). HT ♂ AMS. McAlpine 1960: 79 [illustration of male postabdomen]; Soós 1962c: 449 [key, review, world]; Sasakawa 1971: 57 [New Guinea]. Craspedochaeta delta . Pitkin & Evenhuis 1989: 536 [generic combination, catalogue, Australasia/Pacific].

fascipennis Melander & Argo. View in CoL NT: Tropical Neotropics.

Czernyola fascipennis Melander & Argo 1924: 26 View in CoL . Costa Rica. Cartago Province: La Suiza de Turrialba. HT ♀ USNM. Hennig 1938: 131 [ Bolivia]; Frey 1960: 27 [catalogue, world]; Soós 1962c: 449 [key, review, world], 1968: 3 [catalogue, Americas south of USA].

Craspedochaeta fascipennis . Lonsdale & Marshall 2006b: 55 View Cited Treatment [redescription, generic combination, phylogeny, new records ( Brazil, Costa Rica, Ecuador, Guyana, Mexico, Panama, Peru, Venezuela), illustration of female genitalia].

feminea (Lonsdale & Marshall) . NT: Brazil. NEW COMBINATION

Craspedochaeta feminea Lonsdale & Marshall 2006b: 25 View in CoL View Cited Treatment . Brazil. Amazonas: Manaus, Reserve Ducke. HT ♀ INPA. Lonsdale & Marshall 2006b: 25 View Cited Treatment [phylogeny, illustration of female genitalia].

festiva Soós. View in CoL NT: Brazil (Rio de Janeiro).

Czernyola festiva Soós 1962a: 142 View in CoL . Brazil. Federal District : Rio de Janeiro. HT ♂ USNM. Soós 1962c: 449 [key, review, world], 1968: 3 [catalogue, Americas south of USA] .

Craspedochaeta festiva . Lonsdale & Marshall 2006b: 27 View Cited Treatment [redescription, generic combination, phylogeny, illustration of colouration].

flavofemorata Sasakawa. OR: Japan (Ryukyus), Malaysia (Pen., Sabah) .

Czernyola flavofemorata Sasakawa in Sasakawa & Pong 1990: 58 View in CoL . Malaysia. Sabah: Kinabalu National Park. HT ♂ FRC. Sasakawa & Pong 1990: 58 [illustration of male genitalia, emendation to key in Sasakawa (1971)]; Sueyoshi 2012: 26 [Japan].

Craspedochaeta flavofemorata . Sueyoshi 2006: 12 [generic combination, illustrations of head and male genitalia, photo of wing, Japan], 2011: 9 [ Japan, pictoral key], 2014: 618 [catalogue, Japan]; Sasakawa 2010: 16 [Oriental/Pacific revision, Malaysia].

hyalina Sasakawa. View in CoL AU: Solomon Isl. OR: Malaysia (Sabah).

Czernyola hyalina Sasakawa 1971: 57 View in CoL . Solomon Isl. Tangtalau: Malaita, 200 m. HT ♂ BPBM. Sasakawa 1971: 57 [illustrations of male genitalia].

Craspedochaeta hyalina . Pitkin & Evenhuis 1989: 536 [generic combination, catalogue, Australasia/Pacific]; Sasakawa 2010: 17 [Oriental/Pacific revision, Malaysia, illustration of male genitalia].

loreto (Lonsdale & Marshall) View in CoL . NT: Brazil, Peru. NEW COMBINATION

Craspedochaeta loreto Lonsdale & Marshall 2006b: 71 View in CoL View Cited Treatment . Peru. Loreto Region: Iquitos, 14 km W. HT ♂ USNM. Lonsdale & Marshall 2006b: 71 View Cited Treatment [phylogeny, illustrations of colouration and male genitalia].

maai Sasakawa. OR: Malaysia ( Sarawak). NEW COMBINATION

Craspedochaeta maai Sasakawa 2010: 17 View in CoL . Malaysia. Sarawak: Kapic District , Merirai Village. HT ♀ BPBM [Type No. 17297]. Sasakawa 2010: 17 [illustration of male genitalia].

melanosoma (Lonsdale & Marshall) View in CoL . NT: Brazil, Ecuador. NEW COMBINATION

Craspedochaeta melanosoma Lonsdale & Marshall 2006b: 32 View in CoL View Cited Treatment . Brazil. Amazonas: Reserva Ducke, 26 km NE Manaus. HT ♂ INPA. Lonsdale & Marshall 2006b: 32 View Cited Treatment [phylogeny, illustrations of male genitalia].

nigripalpis (Steyskal & Sasakawa) View in CoL . AU: Indonesia (Papua), Mariana Isl. , Papua New Guinea . OR: Japan (Ryukyus), Vietnam. PA: Japan [Kyushu].

Tonnoiria nigripalpis Steyskal & Sasakawa 1966: 246 . Indonesia. Saipan: Biak Island, Mangrowawa, 50– 100 m. HT ♂ BPBM [Type No. 6727]. Steyskal & Sasakawa 1966: 246 [illustrations of male genitalia].

Czernyola nigripalpis View in CoL . Sasakawa 1971: 58 [generic combination, Biak Island , Papua New Guinea] ; Sueyoshi 2012: 27 [Japan, new record Palaearctic Japan].

Craspedochaeta nigripalpis . Sasakawa 1974: 161 [generic combination, Papua, Papua New Guinea]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific]; Sueyoshi 2006: 12 [illustrations of head and female genitalia, photo of wing, Japan], 2011: 9 [Japan, pictoral key], 2014: 618 [catalogue, Japan]; Sasakawa 2010: 18 [Oriental/Pacific revision, Vietnam].

nigrithorax Hennig. NT: Bolivia, Brazil, Costa Rica, Ecuador, Peru, Trinidad, Venezuela.

Czernyola nigrithorax Hennig 1938: 133 View in CoL . Bolivia. Mapiri: San Carlos, 600m. HT ♂ SMT. Frey 1960: 27 [catalogue, world]; Soós 1962c: 449 [key, review, World], 1968: 4 [catalogue, Americas south of USA]. Craspedochaeta nigrithorax . Lonsdale & Marshall 2006b: 21 [redescription, generic combination, phylogeny, new records ( Brazil, Costa Rica, Ecuador, Peru, Trinidad, Venezuela), illustrations of male and female genitalia].

novaeguineae (Soós) . AU: New Guinea (at least including Papua New Guinea). NEW COMBINATION Chaetoczernyola novaeguineae Sóos 1962a: 141 . New Guinea. “Neu-Guinea”. HT ♀ HNHM. Soós 1962a: 141 [illustrations of head, scutellum and wing]; Sasakawa 1971: 62 [illustrations of male genitalia, Papua New Guinea], 1974: 161 [Bismarck Isl.].

Craspedochaeta novaeguineae . Pitkin & Evenhuis 1989: 536 [generic combination, catalogue, Australasia/Pacific].

oedaleostylis Sasakawa. AU: Bismarck Isl.

Czernyola oedaleostylis Sasakawa 1971: 58 . Papua New Guinea. Manus Island: Momote. HT ♂ BPBM. Sasakawa 1971: 58 [illustrations of male genitalia].

Craspedochaeta oedaleostylis . Sasakawa 1974: 161 [generic combination, Bismarck Isl.]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific].

ornata Hennig. NT: Bolivia, Brazil (São Paulo).

Czernyola ornata Hennig 1938: 132 View in CoL . Bolivia. Mapiri: San Carlos, 800m. HT ♀ SMT. Frey 1960: 27 [catalogue, world]; Soós 1962c: 449 [key, review, world], 1968: 4 [catalogue, Americas south of USA].

Craspedochaeta ornata . Lonsdale & Marshall 2006b: 26 View Cited Treatment [redescription, generic combination, phylogeny, new record ( Brazil), illustrations of colouration and male and female genitalia].

pacaraima (Lonsdale & Marshall) View in CoL . NT: Brazil (Amazonas, Roraima). NEW COMBINATION Craspedochaeta pacaraima Lonsdale & Marshall 2006b: 28 . Brazil. Roraima State: Pacaraima. HT ♂ INPA. Lonsdale & Marshall 2006b: 28 [phylogeny, illustrations of colouration and male genitalia].

palliseta (Malloch) . AU: Society & Marquesas Isl.

Tonnoiria palliseta Malloch 1929a: 99 . French Polynesia. Society Islands: Mo’orea. HT ♂ USNM. Malloch 1932: 215 [on Hibiscus tiliaceus, Hivaoa View in CoL (Marquesas Isl.)].

Czernyola palliseta View in CoL . McAlpine 1960: 79 (as nominal subspecies) [diagnosis]; Frey 1960: 22 [generic combination, summary of original description, catalogue, world]; Soós 1962c: 449 [key, review, world]; Sasakawa 1971: 59 [Society and Marquesas Isl.].

Craspedochaeta palliseta . Pitkin & Evenhuis 1989: 536 [generic combination, catalogue, Australasia/Pacific].

parva Sasakawa. OR: Brunei, Indonesia (Java, Sulawesi), Malaysia (Sabah, Sarawak). NEW COMBINATION Craspedochaeta parva Sasakawa 2010: 18 . Brunei. Belait District, Ulu Belait, Manilas. HT ♂ BPBM [Type No. 17280]. Sasakawa 2010: 18 [illustration of male genitalia].

phaios (Lonsdale & Marshall) View in CoL . NT: Bolivia. NEW COMBINATION

Craspedochaeta phaios Lonsdale & Marshall 2006b: 50 View in CoL View Cited Treatment . Bolivia. La Paz Department: Chulumani, Apa Apa Reserve, 2000 m, 16°21′15ʺS, 68°04′30ʺW. HT ♂ ANCB. Lonsdale & Marshall 2006b: 50 View Cited Treatment [phylogeny, illustrations of male genitalia].

pilosa Sasakawa. OR: Indonesia (Java). NEW COMBINATION

Craspedochaeta pilosa Sasakawa 2010: 19 View in CoL . Indonesia. Java: Tjibodas, 1600m. HT ♂ BPBM [Type No. 17281]. Sasakawa 2010: 19 [illustration of male genitalia].

pleuralis (Williston) View in CoL . NT: Tropical Neotropics*. NEW COMBINATION

Heteroneura pleuralis Williston 1896: 387 View in CoL . St. Vincent. “Leeward side”. HT ♂ BMNH. Czerny 1903: 102 [quotes original description]; Aldrich 1905: 571 [catalogue, North America].

Sobarocephala pleuralis . Melander & Argo 1924: 45 [summary of original description, generic combination, notes on type]; Frey 1960: 27 [catalogue, world]; Soós 1968: 7 [catalogue, Americas south of USA]; Steyskal 1973: 443 [checklist, Antilles].

Craspedochaeta pleuralis . Lonsdale & Marshall 2006b: 44 [generic combination, redescription, phylogeny, new records ( Argentina, Bolivia, Brazil, Costa Rica, Cuba, Colombia, Dominican Republic, Ecuador, El Salvador, Grenada, Mexico, Panama, Peru, Puerto Rico, Trinidad, Venezuela), illustrations of head and male and female genitalia, photo of wing].

pollostos (Lonsdale & Marshall) . NT: Bolivia, Brazil, Costa Rica, Ecuador, Mexico, Panama, Peru, Venezuela. NEW COMBINATION

Craspedochaeta pollostos Lonsdale & Marshall 2006b: 41 View in CoL View Cited Treatment . Costa Rica. Puntarenas Province: R.F. Golfo Dulce, 24 km W Piedras Blancas, 200 m. HT ♂ DEBU. Lonsdale & Marshall 2006b: 41 View Cited Treatment [phylogeny, illustrations of colouration and male and female genitalia].

ponapensis ( Steyskal & Sasakawa, 1966) . AU: Caroline Isl.

Tonnoiria ponapensis Steyskal & Sasakawa 1966: 245 . Federated States of Micronesia: Caroline Islands: Pohnpei Island, Nankep, Mt. Dolen , 540 m. HT ♀ USNM [Type No. 68179].

Czernyola ponapensis View in CoL . Sasakawa 1971: 61 [generic combination, synonymy, diagnosis]. Craspedochaeta ponapensis . Pitkin & Evenhuis 1989: 536 [generic combination, catalogue, Australasia/Pacific].

protomis (Lonsdale & Marshall) View in CoL . NT: Brazil, Costa Rica, Ecuador, Mexico, Peru. NEW COMBINATION Craspedochaeta protomis Lonsdale & Marshall 2006b: 24 . Peru. Loreto Region: Campamento San Jacinto, 175– 215 m. HT ♂ DEBU. Lonsdale & Marshall 2006b: 24 [phylogeny, illustration of female genitalia].

pullipleura (Lonsdale & Marshall) View in CoL . NT: Bolivia.

Craspedochaeta pullipleura Lonsdale & Marshall 2006b: 57 View in CoL View Cited Treatment . Bolivia. La Paz Department: Coroico, 0.5 km SE, 16°11′S, 67°14′W. HT ♂ ANCB. Lonsdale & Marshall 2006b: 57 View Cited Treatment [phylogeny, illustrations of male genitalia, photos of male agonistic behaviour, on dung].

Czernyola pullipleura View in CoL . Lonsdale et al. 2010: 233 [generic combination].

quinquespinula Sasakawa. OR: Philippines. NEW COMBINATION

Craspedochaeta quinquespinula Sasakawa 2010: 21 View in CoL . Philippines. Luzon: Nueva Vizcaya, Dalton Pass. HT ♂ BPBM [Type No. 17282]. Sasakawa 2010: 21 [illustration of male genitalia].

sasakawai (Lonsdale & Marshall) View in CoL . AU: Australia (North Queensland), Bismarck Isl. Caroline Isl., Fiji, Papua New Guinea, Solomons, Vanuatu. OR: Indonesia (Irian Jaya), Malaysia (Sabah), Sri Lanka. NEW COMBINATION Czernyola pleuralis Curran 1936: 54 View in CoL . Solomon Islands. Santa Catalina Island. HT ♂ CAS [Type No. 4067]. Sasakawa 1971: 60 [status reinstated, comparison to palliseta Malloch , Papua New Guinea (previously recorded in Solomons, New Hebrides, North Queensland, Caroline Isl.)]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific]; Sasakawa & Pong 1990: 59 [Sabah].

Czernyola palliseta pleuralis View in CoL . McAlpine 1960: 80 [new status, illustrations of head and surstylus, Queensland]; Soós 1962c: 449 [key, review, world].

Tonnoiria palliseta pleuralis . Steyskal & Sasakawa 1966: 248 [generic combination, Caroline Isl., illustrations of male genitalia].

Craspedochaeta pleuralis . Sasakawa 1974: 162 [generic combination, Bismarck Isl.]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific].

Craspedochaeta sasakawai Lonsdale & Marshall 2006b: 47 View in CoL [replacement for name preoccupied by Williston (1896), new records ( Sri Lanka, Sarawak, Papua New Guinea, Queensland)], 2008b: 5 [redescription, illustrations of egg and male and female genitalia, Fiji]; Sasakawa 2010: 21 [Oriental/Pacific revision, Malaysia].

secunda Sasakawa. OR: Thailand. NEW COMBINATION

Craspedochaeta secunda Sasakawa 2010: 22 View in CoL . Thailand. Pakchong, 100m, NE of Bangkok. HT ♂ BPBM [Type No. 17283]. Sasakawa 2010: 22 [illustration of male genitalia].

sedlaceki Sasakawa. OR: Papua New Guinea.

Czernyola sedlaceki Sasakawa 1971: 61 View in CoL . Papua New Guinea: Morobe Province: Wau. HT ♂ BPBM. Sasakawa 1971: 61 [illustrations of male genitalia].

Craspedochaeta sedlaceki . Pitkin & Evenhuis 1989: 536 [generic combination, catalogue, Australasia/Pacific].

spinulifera Sasakawa. OR: Thailand. NEW COMBINATION

Craspedochaeta spinulifera Sasakawa 2010: 22 View in CoL . Thailand. Chiangmai: Doi Suthep. HT ♂ BPBM [Type No. 17284]. Sasakawa 2010: 22 [illustration of male genitalia].

synneura Hennig. NT: Bolivia, Brazil, Costa Rica, El Salvador, Panama, Trinidad.

Czernyola synneura Hennig 1938: 132 View in CoL . Bolivia. Mapiri: San Carlos. HT ♂ SMT. Frey 1960: 27 [catalogue, world]; Soós 1962c: 449 [key, review, world], 1968: 4 [catalogue, Americas south of USA].

Craspedochaeta synneura . Lonsdale & Marshall 2006b: 48 [generic combination, redescription, phylogeny, new records ( Bolivia, Brazil, Costa Rica, El Salvador, Panama, Trinidad), illustrations of colouration and male and female genitalia (including annulus), photo of wing].

thaiensis Sasakawa. OR: Thailand.

Czernyola thaiensis Sasakawa 1986: 2 View in CoL . Thailand. Nakhon Nayok Province: Khao Yai, ca. 800 m. HT ♂ UOPJ. Sasakawa 1986: 2 [illustration of male genitalia].

transversa (Czerny) . NT: Argentina, Bolivia, Brazil, Costa Rica, Paraguay, Peru.

Craspedochaeta transversa Czerny 1903: 103 View in CoL . “ Peru ”. ST 3♂ 2♀ NMW [location of ST males unknown]. Kertész 1903: 569 [key]; Lonsdale & Marshall 2006b: 39 View Cited Treatment [redescription, phylogeny, new records ( Argentina, Bolivia, Brazil, Costa Rica), illustrations of head and male and female genitalia, photo of wing].

Czernyola transversa View in CoL . Melander & Argo 1924: 27 [generic combination, summary of original description]; Soós 1962c: 449 [key, review, world, new record ( Paraguay)], 1968: 4 [catalogue, Americas south of USA]; Hennig 1938: 131 [ Bolivia]; Frey 1960: 27 [catalogue, world].

trivittata Soós. NT: Brazil.

Czernyola trivittata Soós 1962a: 144 View in CoL . Brazil. Rio de Janeiro State: Terezópolis. HT ♀ USNM . Soós 1962c: 449 [key, review, world], 1968: 4 [catalogue, Americas south of USA].

Craspedochaeta trivittata . Lonsdale & Marshall 2006b: 73 View Cited Treatment [generic combination, redescription, phylogeny, Brazil, illustration of colouration].

unguicauda Sasakawa. OR: Philippines. NEW COMBINATION

Craspedochaeta unguicauda Sasakawa 2010: 24 View in CoL . Philippines. Luzon: Nueva Vizcaya, Dalton Pass, 915m. HT ♂ BPBM [Type No. 17285]. Sasakawa 2010: 24 [illustration of male genitalia].

varicolor (Sueyoshi) View in CoL . OR: Japan ( Ryukyus ). NEW COMBINATION

Craspedochaeta varicolor Sueyoshi 2006: 13 View in CoL . Japan. Ryukyus: Ishigaki Island , Omoto. HT ♂ BLKU . Sueyoshi 2006: 13 [illustrations of head and male genitalia, photo of wing], 2011: 9 [Japan, pictoral key], 2014: 619 [catalogue, Japan].

vietnamensis Sasakawa. OR: Vietnam. NEW COMBINATION

Craspedochaeta vietnamensis Sasakawa 2010: 24 View in CoL . Vietnam. Dilinh (Djiring), 1200m. HT ♂ BPBM [Type No. 17286]. Sasakawa 2010: 24 [illustration of male genitalia].

weemsi (Lonsdale & Marshall) View in CoL . NE: United States (Florida). NT: Jamaica, Mexico, Peru. NEW COMBINATION Craspedochaeta weemsi Lonsdale & Marshall 2006b: 40 . USA. Florida: Dade Co., Everglades National Park. HT ♂ USNM. Lonsdale & Marshall 2006b: 40 [phylogeny, illustrations of colouration and male and female genitalia, photo of wing].

xanthonotum (Lonsdale & Marshall) View in CoL . NT: Brazil, Peru. NEW COMBINATION

Craspedochaeta xanthonotum Lonsdale & Marshall 2006b: 70 View in CoL View Cited Treatment . Peru. Loreto Region: Iquitos (14 km W). HT ♂ USNM. Lonsdale & Marshall 2006b: 70 View Cited Treatment [phylogeny, illustrations of colouration and male genitalia].

xanthopleura Sasakawa. OR: Malaysia (Pen., Sabah). NEW COMBINATION

Craspedochaeta xanthopleura Sasakawa 2010: 26 View in CoL . Malaysia. Sabah: Tarawau District, 19km N of Kalabakan. HT ♂ BPBM [Type No. 17287]. Sasakawa 2010: 26 [illustration of male genitalia].

zongo (Lonsdale & Marshall) View in CoL . NT: Bolivia. NEW COMBINATION

Craspedochaeta zongo Lonsdale & Marshall 2006b: 57 View in CoL View Cited Treatment . Bolivia. La Paz Department: Zongo, 16°06′32ʺS, 68°04′30ʺW. HT ♂ ANCB. Lonsdale & Marshall 2006b: 57 View Cited Treatment [phylogeny].

References

Abbreviations

NEW	University of Newcastle
NMW	Naturhistorisches Museum, Wien
UZMH	University Museum (Zoology)
USNM	Smithsonian Institution, National Museum of Natural History
BPBM	Bishop Museum

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Arthropoda
Class	Insecta
Order	Diptera
Family	Clusiidae

Craspedochaeta hyalina

Sasakawa 2010: 17

Pitkin 1989: 535

Sasakawa 1974: 161

1974

Craspedochaeta caligula

Sasakawa 2010: 16

Pitkin 1989: 536

Sasakawa 1974: 161

1974

Craspedochaeta nigripalpis

Sasakawa 2010: 18

Sueyoshi 2006: 12

Pitkin 1989: 536

Sasakawa 1974: 161

1974

Craspedochaeta oedaleostylis

Pitkin 1989: 536

Sasakawa 1974: 161

1974

Craspedochaeta pleuralis

Pitkin 1989: 536

Sasakawa 1974: 162

1974

Czernyola borneoensis

Sasakawa 1990: 58

Sasakawa 1971: 54

1971

Steyskal 1966: 246

1966

Tonnoiria ponapensis

Steyskal 1966: 245

1966

Tonnoiria palliseta pleuralis

Steyskal 1966: 248

1966

Chaetoczernyola Soós 1962a : 141

Sasakawa 1971: 62

McAlpine 1971: 121

Soos 1964: 6

Soos 1962: 141

1962

Czernyola piceoflava Sóos 1962a : 146

Lonsdale 2006: 59

Steyskal 1973: 443

Soos 1962: 146

Soos 1962: 449

1962

Czernyola festiva Soós 1962a : 142

Soos 1962: 142

Soos 1962: 449

1962

novaeguineae (Soós)

Sasakawa 1971: 62

Soos 1962: 141

1962

Czernyola trivittata Soós 1962a : 144

Soos 1962: 144

Soos 1962: 449

1962

Czernyola basalis brasiliensis Soós 1961 : 399

Lonsdale 2006: 59

Soos 1962: 449

Soos 1961: 399

1961

Czernyola basalis basalis

Soos 1961: 401

1961

Czernyola australis

Pitkin 1989: 535

Soos 1962: 449

McAlpine 1960: 78

1960

Czernyola puncticornis laetior

Sasakawa 1971: 53

Frey 1960: 22

1960

Czernyola bisignata

Pitkin 1989: 535

Sasakawa 1971: 54

Soos 1962: 449

McAlpine 1960: 78

1960

Czernyola boettcheri

Balthazar 1990: 417

Delfinado 1971: 29

Sasakawa 1971: 54

Soos 1962: 449

Frey 1960: 22

1960

Czernyola delta

Pitkin 1989: 536

Sasakawa 1971: 57

Soos 1962: 449

McAlpine 1960: 79

1960

Czernyola palliseta

Sasakawa 1971: 59

Soos 1962: 449

McAlpine 1960: 79

Frey 1960: 22

1960

Czernyola palliseta pleuralis

Soos 1962: 449

McAlpine 1960: 80

1960

Czernyola atrifrons

Pitkin 1989: 535

Sasakawa 1971: 52

Sasakawa 1971: 1971

Steyskal 1966: 245

Soos 1962: 449

Frey 1960: 22

Malloch 1942: 209

1942

Czernyola nigrithorax

Lonsdale 2006: 21

Soos 1962: 449

Frey 1960: 27

Hennig 1938: 133

1938

Czernyola ornata

Soos 1962: 449

Frey 1960: 27

Hennig 1938: 132

1938

Czernyola synneura

Soos 1962: 449

Frey 1960: 27

Hennig 1938: 132

1938

sasakawai

Sasakawa 1990: 59

Pitkin 1989: 536

Sasakawa 1971: 60

Curran 1936: 54

1936

Tonnoiria

Steyskal 1968: 87

Steyskal 1966: 244

Hennig 1952: 227

Malloch 1942: 209

Neave 1940: 508

Malloch 1932: 215

Malloch 1929: 98

1929

Tonnoiria palliseta

Malloch 1932: 215

Malloch 1929: 99

1929

Czernyola puncticornis

Balthazar 1990: 417

Delfinado 1971: 29

Sasakawa 1971: 53

Frey 1928: 106

1928

Czernyola Boettcheri Frey 1928b : 105

Frey 1928: 105

Lonsdale 2006: 52

Steyskal 1973: 443

Soos 1968: 5

Soos 1962: 148

Soos 1962: 449

Soos 1962: 145

Soos 1962: 449

Frey 1960: 27

Melander 1924: 36

1924

Czernyola fascipennis

Soos 1962: 449

Frey 1960: 27

Hennig 1938: 131

Melander 1924: 26

1924

Sobarocephala pleuralis

Steyskal 1973: 443

Soos 1968: 7

Frey 1960: 27

Melander 1924: 45

1924

Czernyola transversa

Soos 1962: 449

Frey 1960: 27

Hennig 1938: 131

Melander 1924: 27

1924

annulipes

Poole 1996: 126

Steyskal 1965: 806

Frey 1960: 27

Brimley 1938: 378

Melander 1924: 29

Malloch 1918: 8

Johnson 1913: 99

1913

Czernyola biseta

Sueyoshi 2012: 26

Balthazar 1990: 417

Sasakawa 1990: 57

Sasakawa 1971: 53

Soos 1962: 449

Frey 1960: 22

Hennig 1941: 146

Hennig 1938: 130

Melander 1924: 26

Hendel 1913: 80

1913

Czernyola

Evenhuis 2016: 49

Sasakawa 1971: 50

Cole 1969: 426

Soos 1962: 449

Frey 1960: 21

McAlpine 1960: 76

Malloch 1942: 209

Neave 1939: 957

Hennig 1938: 130

Curran 1934: 364

Schulze 1928: 912

Melander 1924: 24

Bezzi 1907: 52

1907

Craspedochaeta

Evenhuis 2016: 49

Sasakawa 2010: 13

Lonsdale 2009: 230

Sueyoshi 2006: 12

Papp 2006: 190

Lonsdale 2006: 9

Pitkin 1989: 535

McAlpine 1971: 121

Neave 1939: 860

Schulze 1928: 818

Melander 1924: 24

Czerny 1903: 103

Kertesz 1903: 569

1903

Craspedochaeta basalis

Lonsdale 2006: 64

Czerny 1903: 104

Kertesz 1903: 569

1903

Craspedochaeta transversa

Lonsdale 2006: 39

Czerny 1903: 103

Kertesz 1903: 569

1903

concinna

Lonsdale 2006: 52

Aldrich 1905: 570

Czerny 1903: 101

Kertesz 1903: 570

Williston 1896: 387

1896

Heteroneura pleuralis

Aldrich 1905: 571

Czerny 1903: 102

Williston 1896: 387

1896