Harpalus (Opadius), Casey, 1914

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 692

Subgenus Opadius Casey, 1914

Opadius Casey, 1914: 66. Type species: Cratognathus cordatus LeConte, 1853 by original designation. Etymology. Possibly from the Greek opados (accompanying, attending) [masculine].

Pharalus Casey, 1914: 68. Type species: Pangus testaceus LeConte, 1853 (= Harpalus indianus Csiki, 1932) by monotypy. New synonymy. Etymology. Anagram of the generic name Harpalus [q.v.] [masculine].

Euharpalops Casey, 1924: 116. Type species: Euharpalops wadei Casey, 1924 (= Harpalus fraternus LeConte, 1852) by original designation. New synonymy. Etymology. From the Greek eu (good, agreeable, very), the generic name Harpalus [q.v.], and the suffix - ops (having the appearance of) [masculine].

Haploharpalus Schauberger, 1926: 44. Type species: Harpalus froelichii Sturm, 1818 designated by Habu (1973a: 68). New synonymy. Etymology. From the Greek prefix haplo - (single) and the generic name Harpalus [q.v.] [masculine].

Cordoharpalus Hatch, 1949a: 87. Type species: Harpalus cordifer Notman, 1919 by original designation. Synonymy established by Bousquet and Larochelle (1993: 234). Etymology. From the Latin cordis (heart) and the generic name Harpalus [q.v.] [masculine].

Diversity.

This group is represented in the Nearctic and Palaearctic Regions but the number of species cannot be assessed at this time. The North American fauna has 19 species.

Identification.

The species of this subgenus have been revised by Noonan (1991: 45-127). Lindroth (1968) covered all the species except Harpalus gravis , Harpalus cordatus , and Harpalus apache .

Taxonomic Note.

This taxon corresponds to the fraternus stock of Noonan (1991: 45). Although I have doubt about monophyly of this complex, I have followed Noonan’s (1991) conclusion since it is the last major study on these species. Kataev et al. (2003: 371) listed Opadius Casey and its synonyms as synonyms of the subgenus Harpalus Latreille. In 2010, Kataev included Harpalus apache and Harpalus cordatus (as cordatus group) in the subgenus Glanodes Casey.