Aleiodes (Arcaleiodes) siamensis, Quicke & Butcher, 2012

Butcher, Buntika Areekul, Smith, M. Alex, Sharkey, Mike J. & Quicke, Donald L. J., 2012, A turbo-taxonomic study of Thai Aleiodes (Aleiodes) and Aleiodes (Arcaleiodes) (Hymenoptera: Braconidae: Rogadinae) based largely on COI barcoded specimens, with rapid descriptions of 179 new species, Zootaxa 3457, pp. 1-232 : 18-32

Key to the Aleiodes (Aleiodes) and Aleiodes (Arcaleiodes) species of Thailand

1 Hind trochantellus elongate, dorsal length at level where trochantellus most produced, more than 0.9 times maximum dorsal length of trochanter, and ventral (maximal) length more than 4 x posterior depth; hind coxa strongly transversely striate dorsally ( Fig. 3D ); fore wing 2 nd submarginal cell always short, with vein 3-SR less than 1.2 x length of vein r; hind wing subbasal cell densely and evenly setose; males sometimes with large, setose, lozenge-shaped pits on metasomal tergites 2 and 3; females with large granular appearing gap between widely separated placode sensillae on medioventral part of antenna ( Fig. 4D ); propodeum elongate and angular postero-laterally with posteriorly-directed tooth..................................................................................... Heterogamus (not treated here)

- Hind trochantellus not elongate, dorsal (shortest) length shorter than 0.8 times dorsal length of trochanter, and ventral length less than 4 x posterior depth ( Fig. 182D ); hind coxa usually without oblique striations basally ( Figs 181D ); fore wing 2 nd submarginal cell variable; hind wing subbasal cell variable but often glabrous on at least anterior half; males never with large, setose, lozenge-shaped pits on metasomal tergites 2 and 3; females usually with placode sensillae evenly spaced around median flagellomeres ( Figs 4A,B ) OR if with granular gap ( Fig. 4C ), pronotum not elongate and deep pronope present ( Figs 75D , 141D ); propodeum seldom elongate seldom with postero-lateral angulation with posteriorly-directed tooth (e.g. Fig. 72E ), though may have well-developed tooth on non-elongated propodeum ( Fig. 18E )............... 2

2 (1) Hind wing vein 1-M moderately strongly arched ( Figs 192A , 193A ); fore wing veins 1-M and m-cu distinctly diverging posteriorly ( Fig. 193A ); antennae with white annulus ( Fig. 193A )............................................ 3

- Hind wing vein 1-M moderately more or less straight ( Figs 88G , 97G ); fore wing veins 1-M and m-cu usually distinctly converging posteriorly, rarely more or less parallel ( Fig. 97G ) ( Aleiodes s.s.) OR if diverging, (A. ( Hemigyroneuron )) distal part of fore wing subbasal cell strongly modified and with large glabrous area containing sclerome (see Figs 166D,E ); antennae variable........................................................................... 6

3 (2) Distal part of fore wing subbasal cell strongly modified and often with large glabrous area containing sclerome (see Figs 166D,E ); fore wing subdiscal cell glabrous (see Figs 166B,D,E )............................................................. other S. E. Asian A. ( Hemigyroneuron ) species not yet recorded from Thailand but likely to occur there

- Distal part of fore wing subbasal not modified as above, setose, never with sclerome; fore wing subdiscal cell usually evenly setose [A. ( Arcaleiodes ) species]................................................................. 4

4 (3) Propodeum black ( Fig. 193E ); metasoma striped ( Fig. 193F ), tergite 1 with medial transverse black band, tergite 2 black, with posterior 0.3 white; apex of antenna entirely black; only apical segment of maxillary palp white, basal ones brown or black............................................................................................ 5

- Propodeum pale rufous ( Fig. 192E ); metasoma not striped, tergite 1 orange except cream on apical 0.2, tergite 2 ivory white on basal 0.7, black apically; apex of antenna distinctly whitish ( Figs 192E,F ); two apical segments of maxillary palp white ( Fig. 192C )....................................................... A. (Arcaleiodes) siamensis sp. nov. 5 (4) Fore wing vein 1-CU1 0.6 x 2-CU1; subdiscal cell without longitudinal glabrous line subanteriorly; hind tibia white on basal 0.75, black apically (Fig. 194A); 3 rd metasomal tergite finely longituinally striate on basal 0.75 (Fig. 194F)........................................................................ A. (Arcaleiodes) vanachterbergi sp. nov.

- Fore wing vein 1-CU1 0.75 x 2-CU1; subdiscal cell with longitudinal glabrous line subanteriorly; hind tibia entirely orange ( Fig. 191A ); 3 rd metasomal tergite more strongly longitudinally striate on basal 0.5 ( Fig. 191E )....................................................................................... A. (Arcaleiodes) dresdeni sp. nov.

6 (2) Distal part of fore wing subbasal cell strongly modified and with large glabrous area containing sclerome ( Figs 166D,E ); fore wing subdiscal cell glabrous ( Fig. 165E ); males with glandular pore opening near the midposterior margin of metasomal tergites 4–6........................................................ A. (Hemigyroneron) roberti

- Distal part of fore wing subbasal usually unmodified, rarely largely glabrous BUT never with sclerome; fore wing subdiscal cell usually evenly setose; males [for all those species for which it is known] without gland pore on tergites 4–6 ................................................................................................. 7

7 (6) Fore femur very robust, less than 3.7 x longer from distal part of trochantellus to apex, than maximally deep ( Figs 78A , 116A , 145C ); fore tarsus (excluding aroleum) very short, less than 0.75 x length of fore tibia, its median segment quadrate ( Fig. 145C )....................................................................................... 8

- Fore femur never so robust, always more than 4 x longer from distal part of trochantellus to apex, than maximally deep ( Fig. 121A ); fore tarsus (excluding aroleum) longer, more than 0.82 [minimally 0.85] x length of fore tibia ( Figs 15A , 22A ), its median segment approximately 2 x longer than wide or longer...................................... 11

8 (7) Hind basitarsus strongly laterally compressed and dorsally expanded medially in lateral view, inner face with dense long setae ( Fig. 3A ); size variable but up to 1cm long................................................... Yelicones 1

- Hind basitarsus normal, more or less cylindrical, approximately 4.5 x longer than deep and not dorally swollen medially in profile; small insects with body length approximately 3.5mm [ Aleiodes part]................................... 9

9 (8) Fore femur extremely robust, less than 2.5 x longer from distal part of trochantellus to apex, than maximally deep ( Fig. 78A )......................................................................... A. globifemurus sp. nov.

- Fore femur extremely robust, more than 3 x longer from distal part of trochantellus to apex, than maximally deep ( Figs 116A , 146C ).................................................................................... 10

10 (9) Mesoscutum (except medio-posteriorly) aciculate, rather shiny with some punctures at bases of setae ( Fig. 116C ) and in profile flat-topped and steep anteriorly ( Fig. 116A ); propodeum and metasomal tergites 1–3 largely black ( Figs 116E,F ).................................................................................. A. nonicones sp. nov.

- Mesoscutum coarsely coriaceous ( Fig. 145D ) and dorsally in lateral profile almost continuously curved ( Fig. 145E ); body entirely yellow ( Fig. 145A )........................................................ A. pseudicones sp. nov.

11 (7) Fore wing vein 1-1A giving rise to a distinct, strongly reclivous, anal cross vein sub-medially ( Fig. 165B ); hind wing vein SR with a submedial angulation, usually giving rise to a distinct cross-vein r; fore wing 2 nd submarginal short and trapezoidal, vein 3-SR shorter than vein r ( Fig. 165A )........................................... A. spurivena

- Fore wing vein 1-1A not giving rise to a distinct and strongly reclivous, anal vein medially ( Figs 106D , 127E ); hind wing vein SR without a submedial angulation and without cross-vein r and fore wing 2 nd submarginal more rectangular, vein 3- SR usually longer than vein r........................................................................ 12

12 (11) Distal part of fore wing subbasal cell glabrous ( Fig. 88G ); fore wing subdiscal cell evenly setose ( Fig. 88G ); distal part of fore wing vein M+CU and vein 1-CU1 strongly thickened near intersection with vein 1-M ( Fig. 88G ); largely bright yellow............................................................................ A. kitchingi sp. nov.

- Distal part of fore wing subbasal cell usually evenly setose OR if glabrous THEN EITHER subdiscal cell largely glabrous (especially on anterior half) AND distal part of fore wing vein M+CU and vein 1-CU1 not strongly thickened near intersection with vein 1-M ( Fig. 96E ); colour variable.................................................... 13

13 (12) Basal 3 metasomal tergites forming a carapace with tergite 3 strongly curved in profile concealing all more distal tergites or virtually so AND postero-laterally emarginate ( Figs 32G,H , 59E,F , 181E,F ); mesosternum aciculate; hind coxa strongly transversely to obliquely sculptured basally; small wasps, body length <4.5 mm .............................. 14

- Basal 3 metasomal tergites not forming a carapace concealing all more distal tergites OR IF more or less forming a carapace concealing subsequent tergites ( Fig. 52A ) THEN tergite 3 not posterolaterally emarginate; other characters variable......................................................................................... 16

14 (13) 3rd metasomal tergite approximately 1.2 x longer than 2 nd medially ( Fig. 32G , 59E ); metasoma tergites 2 and 3 at least cream-white laterally, and the 3 rd posteriorly ( Fig. 32H ); 3rd metasomal tergite with postero-lateral corner strongly produced ventrally ( Figs 32H , 59F ); hind wing with vein m-cu distinct ( Fig. 32F ).............................. 15

- 3rd metasomal tergite shorter than 2 nd medially ( Figs 181E ); metasoma tergites 2 and 3 entirely black; 3rd metasomal tergite with posterolateral corner hardly produced ventrally ( Fig. 181F ); hind wing without vein m-cu.... A. trisphaeropyx sp. nov.

15 (14) 1 st metasomal tergite dark posterolaterally, with paler area of variable extent posteromedially ( Figs 59E,F ); 2 nd metasomal tergites largely cream, dark anterolaterally; fore wing vein 3-SR 1.8–1.9 x 2-SR; lateral margin of syntergite when viewed laterally gently curving near level of 2 nd suture ( Fig. 59F ); occipital carina complete mediodorsally, more or less forming a point ( Fig. 59B )................................................................. A. darlingtonia sp. nov.

- 1 st metasomal tergite dark on anterior 0.8 with poterior margin cream coloured all way to lateral corners, with paler area of variable extent posteromedially ( Fig. 32G ); 2 nd metasomal tergites largely black except laterally; fore wing vein 3-SR 1.5 x 2-SR ( Fig. 32F ); lateral margin of syntergite when viewed laterally sharply angled at level of 2 nd suture ( Fig. 32H ); occipital arina broadly incomplete posteromedially ( Fig. 32C )................................ A. butcheri sp. nov.

16 (13) Hind wing vein 2-SC+R transverse and inclivous ( Figs 29D , 87E ) AND relatively large insects with fore wing length> 6.5 mm AND distal part of fore wing subbasal cell largely glabrous ( Figs 29D , 87E ); fore wing uniformly yellow or with apical 0.7 dark, without clear apex; at least hind tarsus largely black ( Fig. 29A )................................ 17

- Hind wing vein 2-SC+R interstitial ( Figs 34D , 109C ) to longitudinal ( Figs 64D , 65E , 82G ) OR if transverse ( Figs 48A , 179E ) THEN smaller insects, less than 6.2 mm fore wing length OR apical part of fore wing subbasal cell largely setose and at most with a small glabrous oval shape surrounded by dense setosity; wing colour variable, sometimes with clear apex ( Fig. 159A ).................................................................................. 18

17 (16) Fore wing venation largely yellow except parastigma and immediately adjoining vein parts black, wing membrane entirely pale yellow-hyaline ( Fig. 29A ).................................................... A. bobwhartoni sp. nov.

- Fore wing venation in distal 0.7 of wing dark brown, wing membrane entirely pale yellow-hyaline on basal third, grey on distal 0.7 ( Fig. 87A ).............................................................. A. jimwhitfieldi sp. nov.

18 (16) Pronotum largely cream white with deep medial pronope usually bordered by a pair of carinate or subcarinate diverging ridges ( Figs 75D , 140C , 141D , 143A ); metasomal tergite 3 strongly curved in dorsal profile, punctate ( Figs 130G ); flagellum often with cream-white band, the flagellomeres in this part of the antenna with large, densely setose gap ventrally, devoid of longitudinal placode sensilla ( Figs. 4C ) [ Aleiodes coronarius group 2]...................... 19

- Pronotum without deep pronope ( Fig. 48A ); metasomal tergite 3 variable, often more or less straight in dorsal profile and often striate ( Figs 63A,F ); flagellum if with cream-white band, not with large gap separating longitudinal placode sensilla in that region (see Figs 4A,B )........................................................................ 45

19 (18) Barcoding sequence known, molecular characters only.................................................... 20 Molecular data unknown, partial morphological key..................................................... 34

20 (19) Bases 1819–1821 coding for V; male antenna with only 1 to 4 apical black flagellomeres beyond white stripe........ 21

- Bases 1819–1821 coding for I; male antenna with more than 8 apical black flagellomeres........................ 23

21 (20) Bases 1657–59 [TTA] coding for L; base 1686 C...................................... A. molecryptus sp. nov.

- Bases 1657–59 [GTA] coding for V; base 1686 T........................................................ 22

22 (21) Bases 1698 1731 1851 1935 2025 A G A T G....................................... A. penultimoluteus sp. nov.

- Bases 1698 1731 1851 1935 2025 G A G C A.......................................... A. apiconigrus sp. nov.

23 (20) Bases 1549–1551 TTG coding for L, 1882–84 GCT coding for A, 1888–90 TTA coding for L... A. concoronarius sp. nov.

- Bases 1549–1551 GTA or GTG coding for V, 1882–84 coding for I, T or V, 1888–90 ATA or ATG coding for M..... 24

24 (23) Bases 1549–51 ATA or ATG coding for M; bases 1873–75 GTA, GTG or GTT coding for V..................... 25

- Bases 1549–51 GTA or GTG coding for V; bases 1873–75 ATT or ATC coding for I............................ 28

25 (24) Bases 1603–05 AGG coding for S; bases 1816 TTA coding for L............................... A. deathi sp. nov.

- Bases 1603–05 GGG coding for G; bases 1816 GTA or GTT coding for V.................................... 26

26 (25) Bases 1807 –09ATT coding for I; bases 1894–96 GTA coding for V; bases 1972 ATA coding for M................................................................................................. A. angulodeum sp. nov.

- Bases 1807 –09GTT coding for V; bases 1894–96 TTA coding for L; bases 1972 TTA coding for L................ 27

27 (26) Bases 1537–39 ACA coding for T;.................................................. A. adorabelleae sp. nov.

- Bases 1537–39 ATA coding for M; ..................................................... A. gaspodei sp. nov.

28 (24) Bases 1946–48 ATT coding for I; bases 1969–71 GCA coding for A......................... A. pronopus sp. nov.

- Bases 1946–48 GTT or GTC coding for V; bases 1969–71 GTA or GCA coding for V or G respecively............. 29

29 (28) Bases 1732–34 TCT coding for S.................................................................... 30

- Bases 1732–34 GCT coding for A.................................................................... 32

30 (29) Bases 1807–09 ACT coding for T; bases 1884–86 ATT coding for I............................. A. cheni sp. nov.

- Bases 1807–09 ATT coding for I; bases 1884–86 GTT coding for V......................................... 31

31 (30) Bases 1591–93 GGA coding for G; bases 1744 GCT coding for A; bases 1816–18 ATA coding for M................................................................................................. A. chenduplus sp. nov.

- Bases 1591–93 AGA coding for S; bases 1744 ATT coding for I; bases 1816–18 GTA coding for V..... A. opus sp. nov.

32 (29) Bases 1816–18 GTA coding for V................................................ A. propodocarinus sp. nov.

- Bases 1816–18 ATA coding for M................................................................... 33

33 (32) Bases 1882–84 GTT coding for V................................................ A. procoronarius sp. nov.

- Bases 1882–84 ACT coding for T.................................................... A. coronopus sp. nov.

34 (19) Propodeum with well-developed posterolateral lobes, with indication of carina [irregular] up posterior edge of lobe ( Figs 15A , 60C ) AND lateral face of lobe coriaceous and distict from sculpture between it and spiracle; male antennae always with more than 8 black flagellomeres beyond white band.................................................. 35

- Propodeum more or less smoothly rounded in profile ( Figs 43A , 46C ) OR IF with some indication of posterolateral lobes

2. The A. coronarius group is one of the more challenging and while molecular data clearly sort them into putative species we have found no good characters to enable all species to be distinguished morphologically. Antennal coloration features appear useful but are sexually dimorphic in some species, and in at least two of the molecularly defined species, there some males show essentially the typical female pattern. Further, the great majority of specimens have one or both antennae broken at or around the white annellar region. Some of the damage looks like they have been bitten. The white zone also appears to have a glandular region of fine pores on one side in a large gap between the placoid sentillae. then lobe with coarse rugulose sculpure laterally continous with sculpture between lobe and spiracle ( Figs 11C , 43E ); male antenna variable, but sometimes with only 1–4 black flagellomeres beyond white band, or entirely black............ 36

35 (34) Propodeum dorsally almost straight with strongly squared posterolateral lobes ( Fig. 15A )....... A. angulodeum sp. nov.

- Propodeum dorsally distinctly curved in profile anteriorly ( Fig. 60C )............................ A. deathi sp. nov.

36 (34) Prododeal sculpture above lateral propodeal carina when viewed laterally, appearing diagonally striate; male antenna with pale stripe occupying only 2 or 3 flagellomeres and separated from apex by at least 15 flagellomeres [pale stripe almost absent in one specimen seen]; female with mesoscutum largely black ( Fig. 46A ), male with dark marks on lateral lobes and anterior of middle lobe of mesoscutum ( Fig. 47D ).................................... A. concoronarius sp. nov.

- Prododeal sculpture above lateral propodeal carina when viewed laterally, not appearing diagonally striate; male antenna with pale stripe variable but sometimes occupying more than 5 flagellomeres.................................. 37

37 (36) Hind coxa basolaterally rather coriaceous............................................... A. pronopus sp. nov.

- Hind coxa basolaterally rather polished with dense punctation............................................. 38

38 (37) Male [unknown for A. propodocarinus sp. nov.]......................................................... 39

- Female [unknown for A. gaspodei , A. molecryptus , A. adorabelleae spp. nov.]................................ 43

39 (38) Black antennal apex beyond white ring comprising only 1 or 2 flagellomeres ( Figs 108A , 130A ) [unknown for A. cheni sp. nov., A. chenduplus sp. nov.]............ A. molecryptus sp. nov., A. penultimoluteus sp. nov., A. apiconigrus sp. nov.

- Black antennal apex beyond white ring comprising more than 10 flagellomeres ( Figs 11D , 143A ) or antenna entirely black ( Fig. 75A )....................................................................................... 40

40 (39) Mesoscutum extensively black anteriorly ( Figs 11D , 75F )................................................. 41

- Mesoscutum entirely rufous dorsally ( Figs 118F , 140C ) sometimes with black lateral margin ( Fig. 143E )............ 42

41 (40) Antenna entirely black ( Fig. 75A )..................................................... A. gaspodei sp. nov.

- Antenna with white ring ( Fig. 11A )................................................. A. adorabelleae sp. nov.

42 (40) White antennal ring occupying more than 6 flagellomeres ( Figs 140A , 143A ) A. procoronarius sp. nov., A. cheni sp. nov.

- White antennal ring occupying fewer than 6 flagellomeres ( Fig. 118A )..... A. opus sp. nov., A. propodocarinus sp. nov.

43 (38) With lateral margin of mesoscutum and anterior of notaulus nearly black, middle lobe of mesoscutum entirely orange ( Fig. 143E ); midlongitudinal carina well developed on anterior 0.75 of propodeum ( Fig. 143E ).... A. propodocarinus sp. nov.

- Mesoscutum entirely orange, or if anteriorly black then also including anterior of middle lobe; midlongitudinal carina well developed on less than anterior 0.65 of propodeum ( Fig. 43F ).............................................. 44

44 (43) Anterior of mesoscutum largely black ( Fig. 43E ); mediodorsal surface of propodeum rather irregularly transversely sculptured ( Fig. 43F ); median area of metanotum rather large and coriaceous dorsally without midlongitudinal carina or deep pit ( Fig. 43F )..................................................................... A. cheni sp. nov.

- Anterior of mesoscutum entirely orange OR with only small dark marks on anterior of each lobe......... other species

45 (18) Hind coxa with transverse to oblique striation on most of basal half, at least on medial side when viewed oblique dorsal angle ( Figs 3C , 36F , 67E , 119B , 157G ) AND occipital carina complete ( Figs 36D , 189D ) and nearly always more or less forming a point mid-anteriorly ( Figs 67C , 119D , 173D ) AND mesopleuron and mesosternum ventrally never smooth and shiny between punctures ( Figs 36E )................................................................... 46

- Hind coxa more or less evenly coriaceous basally (basal to point of maximum width) especially on dorsal and dorsomedial face, not transversely to obliquely sculptured when viewed oblique dorsal angle ( Fig. 156A), though sometimes pimples of aciculation may form parallel oblique lines ( Fig. 3B ), and posterior striation may extend along dorsal edge of dorso-lateral groove of coxa and sometimes more basally in some Chelonorogas group, but in these the mesopleuron and mesosternum are smooth and shiny between punctures; occipital carina variable............................... 66

46 (45) Antenna with a long, rather well-defined white stripe ( Figs 163A , 189A ); back of head black, the mark separated from eye dorsally by paler zone and produced medio-anteriorly to include stemmaticum ( Fig. 189D ); apex of hind tibia with comb of setae (though these not as modified as in many species); hind tarsus conspicuously pale ( Fig. 189A )............. 47

- Antenna without white stripe ( Figs 105A , 157A ); back of head not black; apex of hind tibia with or without comb of setae; hind tarsus dark or same colour as hind tibia............................................................ 49

47 (46) Mesosoma orange-red except pronotum darker submedially ( Fig. 189A ); metasomal tergite 3 strongly arched dorsally in lateral view, and raised above profile of tergite 2 ( Fig. 189A ); metasomal tergites 1–3 black except for pale cream-coloured mark medio-basally on tergite 2, posteriorly ( Fig. 189F ); hind wing vein 2-1A absent............ A. yalaensis sp. nov. Mesosoma black ( Figs 125E, F , 163C ); metasomal tergites 2 and 3 both more or less flat dorsally in profile; metasomal tergites 1 and 2 entirely, and tergite 3 basally, black ( Fig. 125G ), metasomal posteriorly red; hind wing vein 2-1A present as a tubular spur.................................................................................. 48

48 (47) Tarsal claws with 3 large, well-separated pecten spines; fore wing vein 2-CU1 2.2 times 1-CU1 ( Fig. 163F ); 3 rd metasomal tergite with midlongitudinal carina present (weakly) on basal 0.4 ( Fig. 163G ); antenna becoming pale between flagellomeres 16–20 ( Fig. 163A )......................................................... A. s ongsi sp. nov.

- Tarsal claws with pecten formed of line of many adpressed setae (see Fig. 2C ); fore wing vein 2-CU1 1.6 times 1-CU1; 3 rd metasomal tergite with without midlongitudinal carina ( Fig. 125G ); antenna becoming pale between flagellomeres 23–27 ( Fig. 125A )...................................................................... A. parasongsi sp. nov.

49 (46) Hind tibia with comb of especially modified adpressed setae medially and apically (see Fig. 2F ).................. 50

- Hind tibia without comb of especially modified adpressed setae (see Fig. 2E )................................. 52

50 (49) Claws with pecten of approximately 8 short, robust teeth that are longest near the middle of the line and diverge apically ( Fig. 114E ); metasomal tergites almost entirely black except for small ivory spots near mid posterior margins of tergites 1 and 2 ( Fig. 114G )..................................................................... A. niveni sp. nov. - Claws without conspicuous pecten; metasoma not coloured as above........................................ 51

51 (50) Fore wing vein r approximately 0.5 x length of 3-SR ( Fig. 64D ); propodeum and metasomal tergites 1–3 cream medially, with bold black marks laterally ( Fig. 64E,F ); mesopleuron less strongly sculptured posteriorly, more or less aciculate to fine coriaceous posteriorly.......................................................... A. divergerus sp. nov.

- Fore wing vein r approximately 0.8 x length of 3-SR ( Fig. 36A ); propodeum and metasomal tergites 1–3 cream with large bold black marks medially ( Fig. 36E,F ); mesopleuron coarsely sculptured all over, including right up to pleural suture ( Fig. 36E ).............................................................................. A. calicis sp. nov.

52 (49) Fore wing discal cell extremely long and narrow, maximum length more than 2.9 x height at level of 1-M/1-SR+M junction; metasoma orange with tergite margins narrowly darker ( Fig. 67F ); head with black mark behind eye ( Fig. 67D ); maxillary palp smokey grey ( Fig. 67C ); 1st metasomal tergite less than 1.1 x longer than maximally wide ( Fig. 67F ); hind femora basally cream-white, distal 0.7 black or nearly so ( Fig. 67A )......................... A. elongatus sp. nov.

- Fore wing discal cell of normal proportions, maximum length less than 2.7 x height at level of 1-M/1-SR+M junction ( Fig. 157H ); metasomal tergites either strongly patterned black and cream or entirely yellow-brown; maxillary palp white; 1st metasomal tergite usually more than 1.2 x longer than maximally wide; hind femora variable..................... 53

53 (52) 3 rd metasomal tergite distinctly curved dorsally in lateral profile ( Fig. 126F ), usually distinctly narrowing posteriorly from near midlength ( Fig. 126G ), basally finely longitudinally striate becoming acuculate rugulose on posterior third...... 54

- 3 rd metasomal tergite more or less flat dorsally in lateral profile, not or hardly narrowing posteriorly from near midlength, longitudinal striation often coarser and extending nearly to posterior margin of tergite........................... 62

54 (53) Speculum large, smooth and shiny; dorsal part of mesopleuron with rugulose sculpture; metasoma black with broad cream stripe along full length of tergite 2.............................................. A. rectanguliguttatus sp. nov.

- Specular area entirely sculptured, aciculate; dorsal part of mesopleuron with strongly longitudinal pattern of rugae superimposed on coriaceous/rugulose sculpture; metasoma not as above, usually largely yellow brown with transverse brown bands on at least some tergites ( risaae group)...................................................... 55

55 (54) Baso-lateral lobes of 1st metasomal tergite not angularly produced and usually weakly convex posteriorly, not projecting far beyond the small swelling associated with the spiracle though sometimes extending to spiracle ( Fig. 126G ).......................................................................................... A. parisaae sp. nov.

- Baso-lateral flange of 1st metasomal tergite angularly produced and distinctly concave posteriorly and posterior margin of lobe nearly transverse or even reclivous, projecting far beyond the small swelling associated with the spiracle ( Figs 153F , 157I )........................................................................................... 56

56 (55) Body largely orange-red, head largely black except orbits narrowly red, metasomal tergites 3–5 black ( Fig. 157A ); CO1 without amino-acid deletion (codon at positions 2000–2002).............................. A. rugoscutus sp. nov.

- Body brown yellow, head not black, metasoma not patterned as above; CO1 with amino-acid deletion (codon at positions 2000–2002) [not known for A. orion sp. nov.].......................................................... 57

57 (56) Metasoma entirely brown-yellow ( Fig. 105 )............................................. A. mellificus sp. nov.

- Metasoma distinctly patterned black or dark brown on brown-yellow, yellow or cream ( Figs 41A , 91F , 119C , 153F, G ). 58

58 (57) 2 nd metasomal tergite entirely cream white ( Fig. 91F ); 3rd metasomal tergite basally cream white, apically black ( Fig. 91F )................................................................................ A. lemniscus sp. nov.

- 2 nd metasomal tergite variously patterned with black; 3 rd tergite variable...................................... 59

59 (58) 2 nd metasomal tergite largely cream with posterior corners brown, the mark more or less meating at a point medially ( Figs 153F,G ); 3rd metasomal tergite basally yellow-brown, apically black........................... A. risaae sp. nov.

- Metasomal tergite 2 not as above, EITHER black laterally with continuous cream-coloured central band OR at least with dark subtriangular marks extending to anterior corner of tergite; metasomal tergite 3 EITHER black laterally and posteriorly with cream median mark OR virtually entirely black............................................ 60

60 (59) Metasomal tergite 2 black laterally with continuous cream-coloured central band ( Fig. 41E )........ A. centralis sp. nov.

- Metasomal tergite 2 with dark subtriangular lateral marks meeting posteriorly to enclose a lozenge-shaped cream mark ( Figs 119C , 173G )................................................................................. 61

61 (60) Pterostigma ivory-white; mesoscutum with distinct pale brown marks on middle and lateral lobes ( Figs 119B,D ); propodeum largely foveate-rugose anteriorly............................................... A. orion sp. nov.

- Pterostigma dark grey ( Fig. 173F ); mesoscutum uniformly brown-yellow ( Fig. 173E ); propodeum more rugulose, especially antero-medially (Fig. 1734E)................................................. A. synorion sp. nov.

62 (53) Propodeum largely aciculate on either side of midlongitudinal carina, with transverse striae posteriorly ( Figs 10E , 47C ) 63

- Propodeum largely rugulose on either side of midlongitudinal carina ( Fig. 177F )............................... 64

63 (62) Fore wing vein 3-SR 1.5 x 2-SR; 1 st metasomal tergite 1.2 x longer than apically wide ( Fig.47F ); mesoscutum entirely orange ( Fig. 47E ).................................................................... A. conina sp. nov.

- Fore wing vein 3-SR 2.0 x 2-SR; 1 st metasomal tergite 1.9 x longer than apically wide ( Fig. 10E ); mesoscutum with black marks on lateral lobes ( Fig. 10C ).................................................... A. aciculodeum sp. nov.

64 (62) 1st metasomal tergite without or with only weak basal flange that is not sharply angular (see Fig. 10E ); pronotal collar not strongly producted into a neck ( Fig. 176D )............................................................. 65

- 1st metasomal tergite with well developed flange (see Figs 86F ); pronotal collar developed into a long neck ( Fig. 86E ). 66

65 (64) Metasoma black with contiguous cream medial area from middle of tergite 1 to subposterior part of tergite 3 ( Fig. 61F ); flagellum largely black............................................................... A. definus sp. nov.

- Metasomal pattern less clearly defined, pale area brown yellow; mesoscutum entirely pale ochreous yellow ( Fig. 176G ); flagellum orange up to near apex..................................................... A. thirakupti sp. nov.

66 (45, 64) Fourth metasomal tergite strongly rugose to striate sculptured with at least some longitudinal striation ( Figs 23F , 33F , 42G ), posterolaterally angled, in most females and some males with distinct to strong emargination postero-laterally ( Fig. 168F ), always largely exposed and sometimes completely concealing subsequent tergites AND occipital carina broadly interrupted medially ( Figs 23C , 33D , 1910D) AND pronotum not produced into a neck ( Figs 23D ) AND hind tibia without comb of specialised adpressed setae ( Fig. 2E ); fore wing vein 1-CU1 at least 0.7 x length of 2-CU1 ( Figs 34B , 42A , 136A ); 3rd metasomal tergite with complete, or virtually complete mid-longitudinal carina ( Figs 24E , 33F , 83D , 131E ) [ A. buzurae group that form suspended mummies].......................................................... 67

- Fourth metasomal tergite aciculate or smooth and shiny with more or less dense puncturation, often with microsculpture showing distinct transverse pattern, posterolaterally rounded and usually without emargination postero-laterally, OR if with well-developed rugose, rugulose or longitudinally striate sculpture at least on basal 0.5 ( Figs 12A , 50E , 86G , 111F ) THEN occipital carina complete ( Figs 50D , 86C , 111C ) AND / OR pronotum produced into a neck ( Fig. 12C ) OR apex of hind tibia with comb of specialised adpressed setae (see Fig. 2F ) OR fore wing vein 1-CU1 <0.5 x length of vein 2-CU1; 3rd tergite sculpture variable......................................................................... 86

67 (66) Hind wing with vein m-cu distinct ( Fig. 34B ); codon at positions 1795–1797 coding for F; codon at positions 1807–1809 coding for M.................................................................................... 68

- Hind wing without vein m-cu but vein 1-M usually distinctly bent near apex possibly indicating where vein m-cu could originate (Fig, 117A); codon at positions 1795–1797 coding for L; codon at positions 1807–1809 coding for T.......70

68 (67) Third metasomal tergite with dark, sub-circular marks laterally and tergite 4 with large dark mark mediobasally (see Fig. 443D)......................................................................................... 69

- Third and 4th metasomal tergites without dark marks (see Fig. 138E )........................ A. probuzurae sp. nov.

69 (68) Bases 1552–54 CTT coding for L; bases 1570–72 ATT coding for I; bases 1714–16 ATT coding for I; bases 1804–06 ATA coding for M; bases 1948–50 GCA coding for A.................................................. A. buzurae 3

- Bases 1552–54 ATT coding for I; bases 1570–72 GTT coding for V; bases 1714–16 GTA coding for V; bases 1804–06 TTA coding for L; bases 1948–50 TCA coding for S................................ A. buzuriquadruplus sp. nov.

70 (67) First metasomal tergite in lateral view with dorsal carinae uniting near top of tergite, and dorsal profile distinctly depressed medially ( Fig. 175A,C ); flagellum with fewer than 35 (34) segments; pterostigma more or less entirely pale ( Fig. 175A ); posterior margin of 2 nd metasomal tergite more or less evenly curving near midline ( Fig. 175E ). A. tetrarugulosus sp. nov.

- First metasomal tergite in lateral view with dorsal carinae uniting well below top of tergite, and dorsal profile not depressed medially ( Figs 124A , 131A ); flagellum with more than 35 segments; pterostigma with apical 0.3 to 0.5 dark ( Fig. 23A ); posterior margin of 2 nd metasomal tergite more or less straight near midline ( Figs 23F, H ).............. 71

71 (70) Females with posterior margin of 4th metasomal tergite produced into a pair of points sublaterally (at inner border of posterolateral emargination) ( Fig. 23F , 24A,E ) [typical Vietorogas syn. nov. species key here 4]................... 72

- Females with posterior margin of 4 th metasomal tergite not produced into pair of points ( Figs 131E , 168F )........... 75

72 (71) Fore wing vein 1-CU1 1.6 x m-cu.......................................... A. bachma Long & van Achterberg

- Fore wing vein 1-CU1 <1.5 [1.1–1.4] x m-cu.......................................................... 73

73 (72) Antenna pale yellow basally............................................. A. abditivus Long & van Achterberg

- Antenna virtually entirely dark brown, only marginally paler basally......................................... 74

74 (73) Posterior margin of 3rd metasomal tergite rather sharply angled sublaterally; tooth on 4 th tergite sharper ( Fig. 23F )................................................................................... A. bachmaduplus sp. nov.

- Posterior margin of 3rd metasomal tergite not distinctly angled sublaterally; tooth on 4 th tergite more rounded ( Fig. 24E ).............................................................................. A. bachmatriplus sp. nov.

75 (71) Head posteriorly, with region on either side of occipital carina black-brown ( Figs 42C , 131D , 136C ); mesoscutum boldly marked with black ( Figs 42C , 136E )................................................................. 76

- Head posteriorly uniformly pale ( Fig. 80C ); mesoscutum entirely pale cream-white to brown-yellow ( Figs 35D , 80C , 83A )............................................................................................ 81

76 (75) Mesoscutum dark laterally and along line of anterior part of notauli ( Figs 42C , 131C , 168D ); mesopleuron almost entirely dark brown to black; pterostigma entirely black......................................................... 77

- Mesoscutum with dark marks away from edge of each lateral lobe and a pair of sub-medial longitudinal dark marks on the middle lobe ( Figs 98C , 117C ); mesopleuron dark only anterodorsally; pterostigma basally pale ( Figs 98A , 117A )..... 79

77 (76) Face largely ( Fig. 42B ), lateral lobes of mesoscutum largely ( Fig. 42C ) and mesopleuron to below precoxal suture brown to black ( Fig. 42E ); base 1830 T; base 1569 A............................................. A. chamba sp. nov.

- Face largely, lateral lobes of mesoscutum largely and mesopleuron to below precoxal suture pale brown yellow to pale yellow; base 1830 G; base 1569 G.................................................................... 78

78 (77) Bases: 1577 C; 1795 T;1854 T; 1923 T............................................. A. phantasmatis sp. nov.

- Bases: 1577 T; 1795 C; 1854 C; 1923 C............................................ A. subemarginatus sp. nov.

79 (76) Face with submedial brown marks ( Fig. 137B ); tergite 4 with moderately developed semicircular emargination posterolaterally; 1547 A; 1456 A; 1577 T............................................ A. postmaculus sp. nov.

- Face with continuous brown zone across middle; ( Figs 98B , 117B ) tergite 4 hardly emarginate posterolaterally; 1547 not A; 1456 not A; 1577 C............................................................................. 80

80 (79) Bases: 1547 T; 1556 T; 1854 T; 2050 C............................................. A. occimaculatus sp. nov.

- Bases: 1547 G; 1556 C; 1854 C; 2050 T............................................... A. maculiput sp. nov.

81 (75) Pterostigma unicolourous black ( Fig. 35A )........................................... A. buzuritriplus sp. nov.

- Pterostigma bicolorous or largely white ( Figs 80A )...................................................... 82

82 (81) Fore wing vein 1-CU1 as long as or longer than (1.1x) vein 2-CU1.......................................... 83

- Fore wing vein 1-CU1 shorter than vein 2-CU1 ( Fig. 190A )............................................... 84

83 (82) Metasomal tergites 1 and 4 with large nearly black marks, and tergites 2 and 3 with black lateral circular marks ( Fig. 83B–D ); base 1608 C; base 1632 A; base 1736 G................................. A. helenhippersonsae sp. nov.

- Metasoma almost entirely whitish with anterior half of 1 st tergite, and small lateral spots on tergites 2 and 3 dark grey ( Fig. 80D ); base 1608 T; base 1632 G; base 1736 A........................................ A. griseimaculus sp. nov.

84 (82) 1 st tergite 1.2 x wider posteriorly than long; metasoma white with only dark marking on 1 st tergite ( Fig. 124E )........................................................................................ A. parabuzurae sp. nov.

- 1 st tergite less than 1.1 x wider posteriorly than long ( Fig. 190E ); metasoma variable........................... 85

85 (84) Body largely orange-yellow to brown-yellow with extensive black marks on propodeum and metasoma ( Fig. 33 ), dark mark on tergite 1 enclosing a medial pale area ( Fig. 33F )................................ A. buzuriduplus sp. nov.

- Body largely cream-white with small darker marks antero laterally on 1 st metasomal tergite ( Fig. 190 ) A. zuburae sp. nov.

86 (66) Metasomal tergite 4 exposed, with at least basal half strongly sculptured, rugose to longitudinally striate ( Figs 86G ) AND occipital carina complete ( Fig. 86C ) or at most with very narrow medial break AND hind tibia without comb of adpressed setae apico-medially (see Fig. 2E )................................................................... 87

- Metasomal tergite 4 variably exposed but not coarsely sculptured, usually with fine aciculation ( Figs 19E , 38F , 122F , 128E ), and sometimes with these forming a sub-transverse pattern basally ( Figs 61F , 65A ) OR if with some rugulose to striate sculpture basally THEN occipital carina either medially absent ( Figs 30D , 55D ) OR hind tibia with a comb of adpressed setae apicomedially (see Fig, 2F)........................................................... 106

87 (86) Pronotal collar elongate with broad midlongitudinal ridge that is rather clearly demarked posteriorly ( Fig. 140B ); antennae with approximately 15 white segments preceding approximately 4 apical dark ones ( Fig. 140A ). A. procarinatus sp. nov.

- Pronotal collar not elongate OR if elongate ( Figs 72C , 111D ) THEN more or less flat dorsally, without median ridge ( Figs 74D , 111C ); antennae without white stripe............................................................. 88

88 (87) Pronotum elongated with median length> 0.4 x lateral length of mesoscutum measured back to level of posterior corner of pronotum ( Figs 72C , 111D ); hind wing without trace of tubular vein 2-1A; veins 1-1A and cu-a forming a nearly continuous curve ( Fig. 72E ); back of head strongly narrows such that maximum width across occipital carina <0.62 x maximum width of head ( Figs 12E , 68C ) [ Aleiodes narangae group]....................................... 89

- Pronotum not so elongated, with median length <0.4 x lateral length of mesoscutum measured back to level of posterior corner of pronotum ( Figs 100C , 121C , 149E , 182B ); hind wing usually with small but distinct tubular vein 2-1A; veins 1–1A and cu-a forming more of an angle ( Fig. 30G ); back of head variable.................................. 95

89 (88) Body brown extensively marked with black including along side of mesoscutum, propodeum, tergites 1 and 2 basally and laterally, tergite 3 ( Figs 86F ); 3rd and 4th metasomal tergites distinctly emarginate posterolaterally ( Fig. 185B ); codon at bases 1594–96 coding for T......................................................................... 90

- Body almost entirely honey-brown, at most with small darker brown smudge on lateral part of pronotum, small marks medially on 1 st tergite ( A. narangae ) and small submedian brown marks on 2 nd tergite; 3rd and 4th metasomal tergites virtually straight posterolaterally ( Fig. 185A )); codon at bases 88–90 coding for S (unknown for narangae ).........91

90 (89) Metasoma variable, but marked with black and cream-yellow, at least lateral parts of tergites 3 and 4 cream-yellow ( Fig. 86G ); base 1551 A; 1554 T; base1654 T; base 1626 C; base 1731 A; base 1977 T................. A. incisus sp. nov.

- Metasoma orange yellow with dark brown marks ( Fig. 72F ); base 1551 G; base 1554 A; base 1654 C; base 1626 T; base 1731 G; base 1977 C.............................................................. A. fovodeum sp. nov.

91 (89) 1 st metasomal tergite with distinct hour-glass shaped brown mark anteriorly ( Fig. 111E ); 2 nd tergite with small brown lines submedially on anterior half ( Fig. 111E ); midlongitudinal carina of 3 rd tergite well differentiated, rest of tergum rugulose with longitudinal tendency ( Fig. 111F )......................................................... A. narangae

- 1 st and 2 nd metasomal tergites entirely honey-brown ( Figs 68G , 74E ); midlongitudinal carina of 3 rd tergite not or hardly differentiated ( Figs 68G , 74E ), rest of tergum largely with longitudinal striate sculpture......................... 92

92 (91) Codon at postion 1783–1785 coding for V............................................................. 93

- Codon at postion 1783–1785 coding for M............................................................. 94

93 (92) Base 1592 A; base 1679 A; base 1719 T; base 1730 G; base 2082 C; base 2091 G................ A. valinus sp. nov.

- Base 1592 G; base 1679 G; base 1719 C; base 1730 A; base 2082 T; base 2091 A.............. A. etvalinus sp. nov.

94 (92) Base 1592 A; base 1679 G; base 1730 A; base 1836 G; base 1910 G; base 1993 T; base 2008 G; base 2151 A............................................................................................. A. agagg sp. nov.

- Base 1592 G; base 1679 A; base 1730 G; base 1836 A; base 1910A; base 1993 C; base 2008 A; base 2151 T............................................................................................... A. gaga sp. nov.

95 (98) Hind femur short and deep, approximately 4 x longer (measured from narrowing at apex of trochantellus) than maximally deep ( Fig. 121E ); 4th metasomal tergite distinctly though weakly emarginate posterolaterally; metasoma black with posterior of tergite 1 and whole of tergite 2 cream ( Fig. 121E )............................ A. pallimedius sp. nov.

- Hind femur not so swollen, more than 4.2 x longer (measured from narrowing at apex of trochantellus) than maximally deep ( Figs 149A ); 4th metasomal tergite not emarginate posterolaterally; metasoma not coloured as above.......... 96

96 (95) Metasoma entirely black, with striate-rugose sculpture ( Figs 149F , 182F )..................................... 97

- Metasoma not entirely black ( Fig. 100D ), either brown-yellow ( Fig. 38H ), with pale mid-longitudinal area or with pale mid-section; metasomal sculpture variable............................................................. 98

97 (96) 1 st metasomal tergite with more or less straight sides behind distinct protruding basal lobes ( Fig. 149F ); all legs black ( Fig. 149A ); head black, except orbits dorsally narrowly rufous ( Fig. 149D ); body length> 5 mm ... A. reticulisoma sp. nov.

- 1 st metasomal tergite with inwardly-curving sides from base ( Fig. 182F ), without protruding basal lobes ( Fig. 182D ); all legs yellow ( Fig. 182A ); head red except for stemmaticum ( Fig. 182B ); body length approximately 4 mm ... A. tulipus sp. nov.

98 (96) Malar space extremely long such that distance from lowest part of hypostomal carina (viewed laterally) to bottom of eye greater than 0.7 x height of eye in frontal aspect ( Fig. 100B ); fore wing vein 1-M strongly curved ( Fig. 100A ); mesosoma and metasomal tergites 1 and 2 orange, a least tergites 4–6 black ( Figs 100D )................... A. malarius sp. nov.

- Malar space shorter ( Figs 10B , 26B , 30B , 66B ); coloration not as above; other characters variable................ 99

99 (98) Body entirely brown yellow ( Figs 26A , 30A , 66A ).................................................... 100

- Body with prominent black pattern, metasomal tergite 2 black laterally, cream-white medially ( Figs 112F , 188A ).... 102

100 (99) Hind wing vein M+CU 2 x length of 1-M ( Fig. 30G ); fore wing vein r longer than 3-SR ( Fig. 30G ); basal half of metasomal tergite 4 longitudinally striate; apex of fore wing subbasal cell with lozenge-shaped glabrous area ( Fig. 30G ) ................................................................................. A. brevimedius sp. nov.

- Hind wing vein M+CU less than 1.5 [1.1–1.18] x length of 1-M; fore wing vein 3-SR longer than r ( Fig. 26E ); basal half of metasomal tergite 4 variable; apex of fore wing subbasal cell densely setose.................................. 101

101 (100) Dorsal carinae of 1 st tergite forming a sharp triangular point; 1 st tergite approximately 1.6 x longer than wide posteriorly................................................................................ A. basistriatus sp. nov.

- Dorsal carinae of 1 st tergite trilobed, almost square posteriorly; 1 st tergite approximately 1.15 x wider posteriorly than long ( Fig. 66F )........................................................................... A. eadyi sp. nov.

102 (99) Occiput with strong transverse rugulose sculpture ( Fig. 188D ); occipital carina rounded, broken at extreme centre......................................................................................... A. vietuput sp. nov.

- Occiput coriaceous; occipital carina variable......................................................... 103

103 (102) Occipital carina forming an anteriorly directed point medially ( Fig. 112E )..................... A. nathismus sp. nov.

- Occipital carina not forming a point, evenly rounded or forming rounded lobe ( Fig. 21D )....................... 104

104 (103) Propodeum coarsely rugose, short; 1 st metasomal tergite with strong irreglar longitudinal striation ( Fig. 21A )........................................................................................... A. asperum sp. nov.

- Propodeum finely aciculate with only faint rugosity, elongate ( Fig. 10E ); 1 st metasomal tergite strongly aciculate without strong longitudinal striation ( Fig. 10E )............................................................... 105

105 (104) Propodeum evenly aciculate ( Fig. 10E ); lateral lobes of mesoscutum with dark marks ( Fig. 10C ); occipital carina mediodorsally with a distinct posteriorly directed dip........................................ A. aciculodeum sp. nov.

- Propodeum with distinct (though weak) foveate-rugose sculpture on aciculate background (especially when viewed from posterodorsal angle); mesoscutum without dark marks; occipital carina forming a large anterior lobe A. propodealis sp. nov.

106 (86) Mesosternum, mesopleuron and metapleuron densely deeply punctate and or rugose but smooth and shiny between punctures ( Figs 16E , 52C , 76D , 164C ), though occasionally punctures so dense as to be abutting ( Figs 58C , 96C ); typically large insects, fore wing length> 4.5mm; hind wing subbasal cell largely glabrous on anterior half ( Figs 40D , 164C , 171C ); head black, 2 nd and 3rd metasomal tergites never dark laterally and pale medially [species fall in to concept of Chelonorhogas of European authors]................................................................ 107

- Mesosternum, mesopleuron and metapleuron densely aciculate, coriaceous or granular and not shiny between punctures ( Fig. 113D ); often smaller insects; hind wing subbasal cell variable but usually densely and evenly setose ( Fig. 92C ); coloration variable [species fall in to concept of subgenus Aleiodes of European authors]....................... 114

107 (106) Antescutal depression very well-developed, medially occupying more than half length of pronotum in dorsal view ( Fig. 16B ); coxae red, hind tibia basally white, hind tarsus black ( Fig. 16A ); 1 st metasomal tergite short and broad, approximately 0.8 times as long as maximally wide, dorsope normal, with carinae running posteriorly from fused dorsal carinae markedly diverging posteriorly ( Fig. 16D ); propodeum foveate rugose to rugulose anteriorly; fore wing vein 1- CU1 less than 0.5 x length of 2-CU1 ( Fig. 16A )........................................... antescutum sp. nov.

- Antescutal depression narrow and not occupying more than half mid-longitudinal length of pronotum ( Fig. 71D ); IF hind tibia basally white, THEN hind tarsus white ( Fig. 180A ) OR coxae black; 1 st metasomal tergite longer, at least 0.9 times as long as maximally wide ( Fig. 180F ), dorsope variable; other characters variable............................... 108

108 (107) Mid and lower part of mesopleuron and mesosternum smooth and shiny, sparsely punctate, precoxal sulcus virtually absent, indicated only be a very shallow crease dorsally ( Fig. 52C ); mesosoma red, head and metasoma black ( Fig. 52A )......................................................................................... A. convexus

- Mid and lower part of mesopleuron and mesosternum more densely punctate, precoxal sulcus usually deep and narrow but impressed ( Fig. 71C ) OR if less well defined ( Fig. 58C ), then usually indicated by a broad depressed area; not coloured as above ( Figs 58A , 71A , 76A , 96A , 164A , 180A )........................................................ 109

109 (108) Precoxal sulcus not well-defined, whole lower side of mesopleuron very weakly depressed and similarly densely deeply sculptured as adjacent areas with only slight increase in size of punctures in depressed area ( Fig. 58C ); all coxae black ( Fig. 58C ).......................................................................... A. damus sp. nov. - Precoxal sulcus more or less well-defined, usually narrow, depressed and with differentiated, transversely crenulate, coarser sculpture than adjacent areas ( Figs 56D , 71C , 76D , 164C ); all coxae red or yellow ( Figs 56D , 71A , 76D , 164C )....... 110

110 (109) Metasomal tergites 1 and 2 cream-yellow anteriorly, posteriorly black ( Fig. 71D )............ A. flavostriatus sp. nov.

- Metasomal tergites 1 and 2 not striped, either all black or 1st tergite red and sometimes medial part of 2 nd red...... 111

111 (110) Antenna with sharply-defined white anellus (dark grey in one very small male) ( Fig. 76A )........ A. georgiae sp. nov. Antenna entirely black ( Figs 96A , 164A ) or with diffuse paler (yellow-brown) region over broad media zone ( Fig. 180A ).............................................................................................. 112

112 (111) Propodeum and metasomal tergites entirely black ( Fig. 164A ); fore and mid legs and hind femora largely pale yellow...................................................................................... A. sophieae sp. nov.

- Propodeum and at least 1st metasomal tergite red ( Figs 96F , 180F ); fore and mid legs and hind femora orange-red, red or black.......................................................................................... 113

113 (112) Fore wing vein 1-CU1 less than 0.5 x length of 2-CU1 ( Fig. 180E ); flagellum with pale coloured segments over broad medial part ( Fig. 180A ); hind tarsus cream-coloured ( Fig. 180A ); fore wing distinctly darker subapically ( Fig. 180E ); metasomal tergite 3 more convex dorsally in profile ( Fig. 180A )........................... A. tricoloripes sp. nov. Fore wing vein 1-CU1 longer than 2-CU1 ( Fig. 96E ); flagellum unicolorous black ( Fig. 96A ); hind tarsus black; wings entirely hyaline ( Figs 96A,E ); metasomal tergite 3 less convex dorsally in lateral view ( Fig. 96A ). A. longivena sp. nov.

114 (106) Apex of hind tibia without comb of specialised adpressed setae (see Figs 2E , 29E )............................ 115

- Apex of hind tibia with comb of especially modified and adpressed setae on inner side (and usually also ventrally between hind tibial spurs) (see Fig. 2F )...................................................................... 160

115 (114) Occipital carina complete medio-dorsally though sometimes irregular and if so, then [usually] giving rise to at least a faint midlongitdinal carina running to stemmaticum ( Fig. 100C ) [N.B. complete in A. lobocarinus sp. nov. but easily perceived to be medially absent if head not viewed from anterior-dorsal aspect]; size variable........................... 116

- Occipital carina broadly absent medio-dorsally ( Figs 13D , 56D , 103D ); usually small wasps, body length <6 mm .... 147

116 (115) Ocelli enormous almost touching eyes ( Figs 97B , 104D ); claws with strong pecten of dark, discrete teeth.......... 117

- Ocelli of normal size or conspicuously small ( Figs 18C , 100C , 157D ); claws variable but seldom with such strong pecten............................................................................................... 118

117 (116) Hind wing vein 2-SC+R longitudinal ( Fig. 104A ); fore wing vein 3-SR approximately 1.7 x 2-SR ( Fig. 104A ); antennae with slightly less than basal half black, distal part white to tip ( Fig. 104A ); malar space longer ( Fig. 104B )....................................................................................... A. megaophthalmos sp. nov.

- Hind wing vein 2-SC+R quadrate to sub-transverse ( Fig. 97G ); fore wing vein 3-SR less than 1.4 [1.35] x 2-SR ( Fig. 97G ); antennae with black apex ( Fig. 97A ); malar space shorter ( Fig. 97D )................... A. macroophthalmus sp. nov.

118 (116) Malar space extremely long such that distance from lowest part of hypostomal carina (viewed laterally) to bottom of eye greater than 0.7 x height of eye ( Fig. 100B ); fore wing vein 1-M strongly curved ( Fig. 100A ); mandible twisted such that lower tooth directly behind upper one in frontal aspect ( Fig. 100B ); clypeus protruding; mesopleuron with large speculum................................................................................. A. malarius sp. nov.

- Malar space less than 0.65 x height of eye ( Figs 18B , 49B ); other characters variable........................... 119

119 (118) Antenna with a large part, at least from the middle to near apex, pale though not abruptly white-striped [not known for certain in A. conpectenus sp. nov. but probably so] ( Figs 18A , 171A ) AND 2 nd metasomal tergite shorter than posteriorly wide ( Figs 18G , 49F , 171E ); hind tarsus conspicuously cream or white in females ( Figs 18A , 49A ); telotarsi conspicuously dark compared with other tarsal segments ( Fig. 18A ); midlongitudinal propodeal carina complete, running to to semicircular carina just in front of apodeme, though often somewhat irregular on posterior part of propodeum ( Fig. 18F ); ocelli large, distance across ocellar triangle> 4 x shortest distance between posterior ocellus and eye ( Figs 18C , 49D ); moderately large insects, body length> 5 mm .......................................................... 120

- Antenna entirely black ( Figs 155A , 157A ) OR basally red-brown become black towards apex; metasomal tergite 2 proportions, telotarsus colour, hind tarsus colour and propodeal carina variable; ocelli usually smaller, distance across ocellar triangle <4 x shortest distance between posterior ocellus and eye ( Figs 155D , 157D ); size variable......... 122

120 (119) Claws with pecten of approximately 8 long, reddish, clearly separate teeth along whole length.................. 121

- Claws with pecten of approximately 12 adpressed pale teeth along whole length of basal lobe appearing like a silvery or golden lamella at low magnification (see Fig. 2C ) [Highly sexually dimorphic ( Fig. 171A )].. A. subson sp. nov. [females]

121 (120) Mesosoma black with extensive pattern of cream-white markings on mesoscutum, mesopleuron, metapleuron, propodeum ( Fig. 18D E ); metasomal tergites 1 and 2 cream with black marks ( Fig. 18F )................. A. archicolorus sp. nov.

- Mesosoma entirely black (except pale mark on pronotum) ( Fig. 49C, D ); metasomal tergites 1 and 2 entirely red ( Fig. 49F ).............................................................................. A. conpectenus sp. nov.

122 (119) Hind coxa very coarsely sculpured basally both dorsally and medially ( Fig. 157C ); mesoscutum very coarsely sculptured on lateral and anterior of medial lobe ( Fig. 157E )........................................ A. rugoscutus sp. nov

- Hind coxa smooth, aciculate or fine coriaceous basally both dorsally and medially (see Fig. 3B ); mesoscutum not coarsely sculptured on lateral and anterior of medial lobe, generally coriaceous wth more or less dense small punctures at bases of setae ( Figs 14D , 155D )........................................................................... 123

123 (122) Fore wing vein 1-CU1 very short, approximately 0.27 x length of 2-CU1 ( Fig. 155E ); ocelli very small, maximum diameter of posterior ocellus less than 0.75 x shortest distance between posterior ocellus and eye ( Fig. 155D ); antennae very long, approximately 1.5 x wing length ( Fig. 155A ); mesosoma largely black ( Fig. 156A); metasoma red basally, black apically ( Fig. 156B).............................................................. A. rufomedius sp. nov.

- Fore wing vein 1-CU1 longer, at least 0.35 x length of 2-CU1 ( Fig. 133E ); ocelli usually larger, maximum diameter of posterior ocellus more than 0.8 x shortest distance between posterior ocellus and eye ( Fig. 28E ) OR if ocelli small ( Fig. 133D ) THEN antenna shorter OR body entirely yellow ( Fig. 133A )...................................... 124

124 (123) Hind tarsus conspicuously cream or white ( Fig. 28A ); telotarsi conspicuously dark compared with other tarsal segments ( Fig. 28A ); mesoscutum medio-posteriorly cream coloured, otherwise black ( Fig. 28E ); body longer than 5 [6] mm....................................................................................... A. biscutus sp. nov.

- Hind tarsus not conspicuously white, either same colour as hind tibia or yellow or red ( Figs 14A , 40A , 113A ); telotarsi not conspicuously darker than other tarsal articles ( Fig. 40A ); mesoscutum variable; body length variable.............125

125 (124) Hind wing vein M+CU more than 2 [2.15–2.45] x 1-M ( Fig. 133E ); 2 nd submarginal cell very rectangular, vein 3-SR less than 1.5 x 2-SR; basal lobes of tergite 1 small but angularly produced ( Fig. 133F ); metasomal tergites 1 and 2 very strongly and coarsely longitudinally striate ( Figs 14G , 133F )..................................................... 126

- Hind wing vein M+CU less than 2 [1.1–1.8] x 1-M; 2 nd submarginal cell more elongate; basal lobes of tergite 1 and sculpture of 2 nd and 3 rd tergites variable.............................................................. 127

126 (125) Body largely reddish with metasomal tergites 3–6 black ( Fig. 14A,G ); propodeum very coarsely sculptured though without conspicuous mediolateral protuberances ( Fig. 14F )........................................ A. anguaae sp. nov.

- Body entirely pale yellow ( Fig. 133A ); propodeum less coarsely sculptured and with distinct, though weak, posterolateral angulations ( Fig. 133C,D )............................................................. A. placidus sp. nov.

127 (125) Head usually largely black dorsally except orbits more or less narrowly orange red to cream ( Figs 25C , 40E , 113E ) (though with wider orbit mark in A. protocastaneus sp. nov.; Fig. 144D ); metasoma boldly patterned red or orange and black, tergite 2 unicolorous red ( Figs 25E , 40F , 113A ); occipital carina giving rise to at least a faint midlongitudinal ridge running to stemmaticum ( Figs 144D )....................................................................... 128

- Head yellow to red-brown dorsally except for black stemmaticum OR if somewhat darkened medioposteriorly and/or near occipital carina THEN metasoma not boldly patterned red or orange and black, tergite 2 black laterally and pale medially; occipital carina variable........................................................................... 131

128 (127) Midlongitudinal propodeal carina absent on posterior third; female antenna with fewer than 42 [38–40] flagellomeres.129

- Midlongitudinal propodeal carina complete; female antenna with more than 43 [45–47] flagellomeres.............. 130

129 (128) Metasomal tergite 1 entirely red ( Fig. 113F ); metasomal tergites 4–7 black ( Fig. 113F ); back and top of head largely black dorsally with sharply defined brown-yellow orbits ( Fig. 113E ).............................. A. necsubson sp. nov.

- Metasomal tergite 1 black mark anteriorly enclosing yellow semicircle posteriorly ( Fig. 144E ); metasomal tergites 4–7 entirely orange yellow ( Fig. 144E ); back and centre of top of head dark brown black, yellowish orbits wider and less clearly defined ( Fig. 144D )...................................................... A. protocastaneus sp. nov.

130 (128) Second metasomal tergite 1.1 x wider posteriorly than long, and 1.1 x wider han anteriorly wide ( Fig. 40F ); codon at bases 1594–96 coding for S; codon at bases 1597–99 coding for G; codon at bases 1816–18 coding for V; codon at bases 1990–93 coding for M............................................................. A. castaneus sp. nov.

- Second metasomal tergite parallel-sided and longer than wide ( Fig. 25E ); codon at bases 1594–96 coding for T; codon at bases 1597–99 coding for S; codon at bases 1816–18 coding for M; codon at bases 1990–93 coding for T.......................................................................................... A. basipunctatus sp. nov.

131 (127) Metasoma entirely yellow to brown yellow, at most slightly darkened anteriorly, without obvious median pale line ( Figs 26A , 66A )...................................................................................... 132

- Metasoma dark with a pale median line at least on tergite 2 ( Figs 57F , 148E , 178E )............................ 137

132 (131) Propodeum with strong mediolateral protuberances ( Figs 65C,F ); setosity of distal part of fore wing membrane approximately twice as dense as that of basal cell ( Fig. 65E )................................. A. downeyi sp. nov.

- Propodeum without strong mediolateral protuberances ( Figs 26C , 50C , 54E ); setosity of distal part of fore wing membrane approximately same density as that of basal cell........................................................ 133

133 (132) Body more than 5.5 [6.0-6.6] mm long; 1 st metasomal tergite approximately 1.5 x longer than posteriorly wide; basolateral lobes of 1 st metasomal tergite distinctly protruding, rather angular ( Fig. 50E )................................ 134

- Smaller, body less than 5.5 [3.8-5] mm long; 1 st metasomal tergite approximately 1.1 x wider posteriorly than long; basolateral lobes of 1 st metasomal tergite variable, hardly ( Fig. 120E ) to moderately produced, rounded and posteriorly convex.......................................................................................................................................................................................... 135

134 (133) Occiput evenly aciculate without transverse sculpture ( Fig. 26D ); antennae brown yellow basally becoming black from near the middle ( Fig. 26A )......................................................... A. basistriatus sp. nov.

- Occiput with distinct transverse corruscation ( Fig. 50D ); antennae brown yellow on basal 0.75 becoming black apically ( Fig. 50A )...................................................................... A. constriatum sp. nov.

135 (133) Occiput rather coarsely sculptured with distinct transverse striation superimposed on coriaceous-punctate sculpture ( Fig. 54D ); body almost entirely brown-orange, metanotum darker ( Fig. 54A ); wing membrane brownish and conspicuously darker near veins; 2 nd metasomal tergite 1.2 x wider posteriorly than long ( Fig. 54F ); antenna basally red-brown become black towards apex ( Fig. 54A ); antenna with approximately 38 flagellomeres................. A. corrusciput sp. nov.

- Occiput coreacous with hardly any indication of transverse sculpture ( Figs 110B , 120B ); body pale yellow, metanotum not conspicously darker ( Fig. 110D ); wing membrane clear to pale grey ( Fig. 110A ); antenna with approximately 41 or more flagellomeres [not known for certain in A. mutilus sp. nov.].............................................. 136

136 (135) Occipital carina not or hardly complete mediodorsally where it is weak and rather pointed and confused with adjacent sculpture ( Fig. 120B ); with dark grey marks on pronotum laterally, median part of propodeum and basal and anterolateral part of 1 st metasomal tergite ( Fig. 120A,C,E )......................................... A. paenicarinus sp. nov. - Occipital carina strong and complete medio-dorsally, more or less evenly rounded ( Fig. 110D ); body entirely pale yellow ( Fig. 110A )......................................................................... A. mutilus sp. nov.

137 (131) Ovipositor and ovipositor sheaths strongly upcurved ( Fig. 57A )........................... A. curvicauda sp. nov.

- Ovipositor and ovipositor sheaths not strongly upcurved ( Figs. 127A , 178A )................................ 138

138 (137) Metasomal tergites 1–4 black except for a parallel-sided ivory white mark down middle of tergite 2 and occasionally a very small pale area midposteriorly on tergite 1 ( Figs 148E , 178E )............................................. 139

- Metasoma with more extensive pale marks, at least antero-medial part of tergite 4 with large pale mark ( Figs 127F ).. 140

139 (138) Antenna with fewer than 38 [35] flagellomeres; propodeum rather strongly arched in dorsal profile ( Fig. 148D ); hind femur, tibia and tarsus largely brown yellow ( Fig. 148A ); claws with pecten of 7 or 8 long, fine, pale brown teeth on basal lobe, these splaying and distinctly separated, and the penultimate ones slightly longer (see Fig. 2D )............................................................................................ A. rectanguliguttatus sp. nov.

- Antenna with more than 39 [41] flagellomeres; propodeum shallowly arched in dorsal profile ( Fig. 178C ); hind femur, tibia and tarsus largely black ( Fig. 178A ); claws with at most a few small pecten teeth on basal part of basal lobe, not conspicuous........................................................................ A. tobiasi sp. nov.

140 (138) Midlongitudinal propodeal carina not extending to posterior or propodeum, replaced by transverse crenulae; lateral margin of propodeum rufous, mesosoma without distinct dark line along side ( Fig. 127C ); fore wing vein 1-CU1 0.55 x 2-CU1 ( Fig. 127E ); largely orange yellow, without dark marks on back of head ( Fig. 127D ), lateral parts of thorax or lateral part of propodeum more or less uniformly honey yellow without conspicuous dark line ( Fig. 127A,C )... A. paulmarshi sp. nov.

- Midlongitudinal propodeal carina extending to posterior or propodeum though sometimes irregular posteriorly ( Fig. 47F ); lateral margin of propodeum black, mesosoma with distinct dark line along side; fore wing vein 1-CU1 usually less than x 2-CU1; coloration variable......................................................................... 141

141 (140) Hind femur and tibia patterned, at least apical 0.25 of hind femur dark brown to black ( Figs 47A , 186A )........... 142

- Hind femur and tibia uniformly pale coloured, whitish, yellow or orange yellow ( Figs 81A , 82A , 95A )............ 143

142 (141) Hind femur whitish basally with apical 0.7 dark yellow-brown, hind tibia distinctly banded with narrow white ring near base, followed by two broad dark brown-yellow ( Fig. 186A ); mesoscutum brown with dark marks on each lateral lobe and on medial lobe ( Fig. 186F )........................................................ A. variifemurus sp. nov.

- Hind femur more or less uniformly brown beyond pale base ( Figs 47A ); mesoscutum largely orange brown ( Fig. 47E )...................................................................................... A. conina sp. nov.

143 (141) Mesoscutum with extensive dark markings, nearly black along line of notauli ( Fig. 81D ); face with dark medial mark ( Fig. 81B ); legs rather robust with tarsal articles markedly deepening towards apex ( Fig. 81A ); antenna with more than 37 [41] flagellomeres; body more than 5 [5.8] mm long............................................ A. guidaae sp. nov.

- Mesoscutum and face uniformly yellow or brown yellow ( Fig. 147B,D ) OR if with dark marks, the lines of the notauli not dark ( Fig. 82C ); legs rather less robust ( Fig. 82A ); antenna with fewer than 36 [27–35] flagellomeres; body less than 4.5 mm long....................................................................................... 144

144 (143) Body largely brown yellow with dark brown to black mark on lateral lobes of mesoscutum and metapleuron ( Figs 82A,C )....................................................................................... A. hei sp. nov.

- Body largely cream white with dark brown line laterally including mark on lateral part of back of head, side of pronotum, upper part of mesopleuron and lateral part of propodeum ( Figs 95A,C )...................................... 145

145 (144) Occipital carina strongly arched medio-dorsally into a lobe where it is rather weakly defined ( Fig. 95C ); fore wing vein 1- CU1 less than 0.44 x 2-CU1; antenna with fewer than 29 [27] flagellomeres................. A. lobocarinus sp. nov.

- Occipital carina evenly rounded medio-dorsally or forming a point, not formed into a lobe; fore wing vein 1-CU1 more than 0.45 [0.46–0.5] x 2-CU1; antenna with more than 30 [31–34] flagellomeres.............................. 146

146 (145) Metasomal tergite 4 brown on anterior 0.7 ( Fig. 147F ); occiput coriaceous ( Fig. 147D ); head brown posteriorly but without distinct mark dorsolaterally on occipital carina ( Fig. 147D )................................... A. ptraci sp. nov.

- Metasomal tergite 4 entirely white ( Fig. 79F ); occiput aciculate ( Fig. 79D ); head with small dark mark next to occipital carina dorsolaterally ( Fig. 79D )........................................................ A. glutinum sp. nov.

147 (115) Claws strongly pectenate with 12 or more strong, diverging teeth running from base of basal lobe to base of claw proper; body length 5.5–7.2 mm; females with antenna with large pale band medially and metasomal tergites 3–6 black ( Fig. 171A , right)........................................................................ A. subson sp. nov.

- Claws without conspicuous pecten; smaller wasps; coloration of female not as above......................... 148

148 (147) Metasoma boldly patterned with dark marks ( Figs 19C , 57E, G , 103E , 150C )................................. 149

- Metasoma largely white, without dark markings OR at most with weak grey markings on base and sides of 1 st tergite ( Figs 13E , 44C , 169D )................................................................................ 152

149 (148) Hind coxa largely dark brown to black ( Fig. 19A ); midlongitudinal propodeal carina incomplete, absent on posterior 0.25................................................................................. A. areelucki sp. nov.

- Hind coxa entirely brown yellow ( Figs 103E , 150C ); midlongitudinal propodeal carina complete though may be slighly irregular posteriorly.............................................................................. 150

150 (149) Ovipositor and ovipositor sheaths strongly upcurved ( Fig. 57A,F ); metastoma with tergite 3 brown yellow, darker medially ( Fig. 57F ); pterostigma largely white with narrow grey brown border ( Fig. 57A )............... A. curvicauda sp. nov.

- Ovipositor and sheaths straight ( Fig. 103A ); metasoma not as above, tergite 3 darker laterally ( Figs 103G , 150E ); pterostigma with distal 0.5 dark ( Fig. 103A )........................................................... 151

151 (150) Malar space (distance from lowest part of hypostomal carina to bottom of eye, in frontal aspect) more than 0.6 [0.7] x height of eye ( Fig. 150B ); speculum well developed ( Fig. 150C ); mesosoma largely cream white with dark brown black lateral stripe, metanotum and propodeum (Fig,. 150C)......................................... A. reus sp. nov.

- Malar space (distance from lowest part of hypostomal carina to bottom of eye, in frontal aspect) less than 0.6 [0.5] x height of eye ( Fig. 103B ); speculum absent, specular area entirely aciculate ( Fig. 103E ); mesosoma largely orange yellow with dark brown to black marks on side of pronotum, metanotum and propodeum ( Fig. 103E )..... A. mediomaculatus sp. nov.

152 (148) Fore wing vein 3-SR shorter than vein r ( Fig. 57A )....................................... A. curtivena sp. nov.

- Fore wing vein 3-SR at least 1.2 x longer than vein r ( Figs 13A , 44D )....................................... 153

153 (152) Frons with depressions behind antennal sockets distinctly differentiated from rest of frons and with at least a weak “V” or “Y”-shaped carina in front of anterior ocellus ( Figs 44B , 162D ); ocelli usually relatively large, the maximum diameter of posterior ocellus often greater than shortest distance between posterior ocellus and eye ( Figs 13D , 44B ); pterostigma with apical half dark grey ( Fig. 44D ); propodeum and 1 st metasomal tergite completely yellow or brown yellow ( Fig. 44A ) 154

- Frons more or less flat ( Fig. 170D ) or if with distinct depressions behind antennal sockets then these merging gradually with rest of frons and without “V” or “Y”-shaped carina in front of anterior ocellus; ocelli sometimes with maximum diameter of posterior ocellus less than 0.8 than shortest distance between posterior ocellus and eye (Figs); pterostigma usually completely pale ( Figs 73A , 169A ); propodeum and base of 1 st metasomal tergite usually marked with grey brown ( Fig. 73E )..... 156

154 (153) Codon at 1543–45 coding for A; codon at 1582–84 coding for L; codon at 1594–96 coding for S; codon at 1597–99 coding for S; codon at 1720–23 coding for L; codon at 1807–09 coding for I; codon at 1816–18 coding for V................................................................................................ A. alboluteus sp. nov.

- Codon at 1543–45 coding for S; codon at 1582–84 coding for M; codon at 1594–96 coding for T; codon at 1597–99 coding for G; codon at 1720–23 coding for M; codon at 1807–09 coding for M; codon at 1816–18 coding for I............ 155

155 (154) Codon at 1888–1890 coding for A....................................................... A. codon sp. nov.

- Codon at 1888–1890 coding for T.................................................. A. similicodon sp. nov.

156 (153) Codon at 1732–1734 coding for A; codon at 1795–1797 coding for M; codon at 1816–1818 coding for L; codon at 1819–1821 coding for T; codon at 1884–1886 coding for P; codon at 1900–1902 coding for T.. A. fuscomedius sp. nov.

- Codon at 1732–1734 coding for G or S; codon at 1795–1797 coding for T or I; codon at 1816–1818 coding for I; codon at 1819–1821 coding for I; codon at 1884–1886 coding for S; codon at 1900–1902 coding for M................... 157

157 (156) Codon at 1732–1834 coding for S................................................ A. subfuscomedius sp. nov.

- Codon at 1732–1834 coding for G................................................................... 158

158 (157) Codon at 1795–1797 coding for I................................................... A. paenicarinus sp. nov.

- Codon at 1795–1797 coding for T................................................................... 159

159 (158) Base at 1751 A; base at 1814 G; base at 1929 T........................................ A. mediofuscus sp. nov.

- Base at 1751 G; base at 1814 A; base at 1929 C..................................... A. submediofuscus sp. nov.

160 (114) Fore wing vein r> 1.25 x length of vein 3-SR ( Fig. 30G , 187E ) AND fore wing vein 1-1A medially with distinctly thickened and curved zone ( Fig. 30G , 187E ); occipital carina medially obliterated ( Figs 30D ); ocelli large, distance across ocellar triangle 7 or 8 x shortest distance between posterior ocellus and eye ( Figs 30D , 55D , 177D ); hind wing vein M+CU more than 1.9 x length of 1-M ( Fig. 30G ); basal lobes of 1 st metasomal tergite sharply angularly projecting ( Fig. 115E ).................................................................................................. 161

- Fore wing vein r <1.15 x length of vein 3-SR ( Figs 48E , 107A ); fore wing vein 1-1A without markedly thickened and curved zone medially ( Fig. 106D ); occipital carina, ocelli and hind wing vein M+CU and 1 st tergite variable....... 166

161 (160) 4 th metasomal tergite exposed and distinctly longitudinally striate on basal half medially ( Fig. 55F ); speculum absent, entie aea aciculate.................................................................................... 162

- 4 th metasomal tergite smooth or aciculate (if exposed) without longitudinal striation; speculum variable............ 164

162 (161) Occiput aciculate with quite deep punctures associated with setae; midlongitudinal carina of propodeum complete; colour largely brown yellow ( Fig. 30A ) ..................................................................... 163

- Occiput aciculate-coriaceous ( Fig. 55D ); midlongitudinal carina of propodeum absent posteriorly ( Fig. 55E ); colour largely cream white, centre of propodeum pale brown ( Fig. 55E ).............................. A. cramum sp. nov.

163 (162) Hind wing vein M+CU less than 2.0 [1.9] x 1-M ............................................ A. nivori sp. nov.

- Hind wing vein M+CU more than 2.0 [2.2] x 1-M....................................... A. brevimedius sp. nov.

164 (161) Fore wing vein 1-CU1 0.85 x as long as vein 2-CU1....................................... A. tmaliaae sp. nov.

- Fore wing vein 1-CU1 less than 0.8 x as long as vein 2-CU1 ( Fig. 177A ).................................... 165

165 (164) Fore wing vein 1-CU1 less than half length of vein 2-CU1 ( Fig. 90A ); codon at positions 1795–97 coding for M; codon at positions 367–9 coding for M...................................................... A. lavaeolous sp. nov.

- Fore wing vein 1-CU1 more than half length of vein 2-CU1 ( Fig. 187E ); codon at positions 1795–97 coding for T; codon at positions 367–9 coding for I........................................................ A. vetinarii sp. nov.

166 (160) Tergite 3 entirely smooth and shiny ( Fig. 134G ); legs robust............................... A. polititergus sp. nov.

- Tergite 3 at least coriaceous, usually rugulose or longutudinally striate at least on basal third.................... 167

167 (166) Ocelli minute and ocellar triangle very small, transverse diameter of posterior ocellus less than 0.3 x distance between stemmaticum and occipital carina, and distance across ocellar triangle less than 0.7 (0.68) distance between stemmaticum and occipital carina ( Fig. 107C ); fore wing vein 1-CU1 0.8–1.1 x 2-CU1; basal 3 metasaomal tergites dark brown to black.......................................................................... A. microophthalmos sp. nov.

- Ocelli not minute and ocellar triangle larger, transverse diameter of posterior ocellus more than 0.35 times distance between stemmaticum and occipital carina and/or distance across ocellar triangle more than distance between stemmaticum and occipital carina; fore wing vein 1-CU1 shorter than 2-CU1 OR IF rarely up to 1.1 x 2-CU1 THEN ocelli normal sized OR occiput with distinct wide median furrow; metasoma differently coloured..................... 168

168 (167) Distance across stemmaticum approximately equal to shortest distance between eyes; fore wing grey medially with clear apex ( Fig. 159A ); antennae largely orange yellow becoming dark towards tip ( Fig. 159A )......... A. scottshawi sp nov.

- Distance across stemmaticum less than 0.9 x shortest distance between eyes; forewing not as above, without clear apex ( Figs 37D , 106B ); antenna usually largely black, sometimes with white apex ( Fig. 37A )........................ 169

169 (168) Occipital carina complete medio-dorsally ( Figs 37D , 48B , 77B ), very rarely in species where carina produced anteriorly to form a point, extreme centre may be weak or incomplete ( Fig. 64C )........................................ 170

- Occipital carina distinctly incomplete medio-dorsally, curving anteriorly near level of posterior ocelli and becoming obliterated, line seldom demarked by clear transition in sculpture ( Figs 99B )................................ 199

170 (169) Hind wing vein 2-SC+R distinctly transverse ( Figs 37F , 48A ) AND hind wing subbasal cell entirely glabrous ( Figs 37G , 48F ) AND antenna with white apex ( Fig. 37A ) [not known in glabribasalis sp. nov.]; fore wing 1st subdiscal cell with distinct longitudinal glabrous zone in distal part of cell.................................................. 171

- Hind wing vein 2-SC+R longitudinal or interstitial OR if transverse THEN hind wing subbasal cell almost entirely setose ( Fig. 106C ); fore wing 1st subdiscal cell usually without distinct longitudinal glabrous zone in distal part of cell; antenna and mesonotum variable.......................................................................... 173

171 (170) Mesosoma largely honey-yellow dorsally and ivory to cream ventrally with black lateral line running from side of pronotum, upper part of mesopleuron and side of propodeum ( Fig. 48A,D ).................. A. connudatum sp. nov.

- Mesosoma at least with lateral lobes and anterior of medial lobe of mesoscutum and most of metanotum and propodeum black.......................................................................................... 172

172 (171) Scutellum and posterior posterior of middle lobe of mesonotum cream ( Fig. 77D ); metasomal tergites 1–3 black with triangular cream marks posteriorly on tergites 1 and 2 ( Fig. 77F )........................ A. glabribasalis sp. nov.

- Scutellum and mesonotum entirely black ( Fig. 37C,F ); metasomal tergites 1–3 black with with only small cream mark posteriorly on tergite 2 ( Fig. 37E )........................................................ A. calvus sp. nov.

173 (170) Tarsal claws with pecten composed of line of approximately 10–15, parallel, adpressed teeth giving appearance of a silvery lamella ( Fig. 2C )................................................................................. 174

- Tarsal claws without conspicuous pecten OR if with pecten THEN it is composed of 3 –7 widely separated teeth (see Fig. 2B ) OR composed of 7–10 longer, setiform teeth that diverge, the longer being subapical ( Fig. 2D ).............. 180

174 (173) Occipital carina produced mid-anteriorly to form a point and giving rise to a short carina ( Fig. 129B,F ); codon at position 1831–1833 coding for I.......................................................................... 175

- Occipital carina evenly rounded anteriorly ( Fig. 132D ) or if pointed, not giving rise to a short midlongitudinal carina ( Figs 128C ); codon at position 1831–1833 coding for V..................................................... 176

175 (174) Tergite 3 distinctly finely longitudinally striate on anterior 0.4 ( Fig. 129C ); tergites 1 and 2 anteriorly dark grey-brown ( Fig. 129C ); bases 1704, 2025, 2070 G, G, C respectively................................. A. pectunguis sp. nov.

- Tergite 3 hardly striate on anterior 0.4 ( Fig. 129G ); tergites 1 and 2 anteriorly largely pale ( Fig. 129G ); bases 1704, 2025, 2070 A, A, T respectively....................................................... A. pectunguisella sp. nov.

176 (174) Mesosoma largely cream-white or brown yellow with or without black line running along upper part of pronotum and mesopleuron and along side of propodeum ( Figs 106E , 132C )............................................ 177

- Mesosoma either largely black or largely orange-red ( Figs 38C )........................................... 179

177 (176) Mesosoma entirely cream-white to yellow; metasomal tergite 5 without any dark mark, palest of the three species in this complex ( Fig. 132 )................................................................. A. pinnulae sp. nov.

- Mesosoma largely cream-white with black line running along upper part of pronotum and mesopleuron and along side of propodeum ( Figs 22A , 106E )...................................................................... 178

178 (177) Metasomal tergite 3 with a pair of lateral black marks ( Fig. 22E ); propodeum without midlongitudinal carina, with weak smooth medial line on anterior 0.2 only ( Fig. 22F ); base 1656 G; base 1854 T; base 1894 A........... A. atuin sp. nov.

- Metasomal tergite 3 with a single large transverse black mark ( Fig. 106F ); propodeum with distinct midlongitudinal carina on basal 0.6 ( Fig. 106C ); base 1656 A; base 1854 C; base 1894 G............................. A. mericeti sp. nov.

179 (176) Flagellum black basally becoming white from approximately the midpoint ( Fig. 38A ); mesoscutum with lateral lobes and anterior of middle lobe largely black, posteromedially cream ( Fig. 38D ); codon at position 277–9 coding for M; posterior of metasomal tergite 3 and all of tergite 4 orange............................................. A. canus sp. nov.

- Flagellum entirely black or basally yellow-brown becoming dark or black towards apex ( Fig. 128A ); mesoscutum largely orange-red with propodeum somewhat darker; codon at position 277–9 coding for I; metasomal tergites 3 and 4 entirely black ( Fig. 128F )................................................................. A. pectopulicis sp. nov.

180 (173) Fore tibia long and curved, more than 1.5 x length of fore tarsus ( Figs 122A , 123A ); pronotum laterally with coarse radially arranged rugae medially and posteriorly ( Fig. 123E ); 3 rd metasomal tergite less transverse ( Figs 122F , 123G ) 181

- Fore tibia not especially elongate and more or less straight, less than 1.5 x length of fore tarsus ( Fig. 137A ); pronotum more or less coriaceous posterolaterally without strong radial or parallel pattern of rugae ( Fig. 137F ); 3 rd metasomal tergite more transverse ( Fig. 137E )........................................................................ 182

181 (180) Base 1571 A; base 1625 C; base 1637 G; base 1694 C; base 1733 G........................... A. palmae sp. nov.

- Base 1571 G; base 1625 T; base 1637 A; base 1694 T; base 1733 C........................ A. palmatipes sp. nov.

182 (180) Antenna with more than 49 flagellomeres penultimate flagellomeres (those approximately 10 from apex) approximately 1.5–2 x longer than wide ( Fig. 4B )................................................................. 183

- Antenna with fewer than 48 flagellomeres OR if with approximately 50 flagellomeres THEN penultimate flagellomeres (those approximately 10 from apex approximately 2.5–3 x longer than wide ( Fig. 4A )........................ 186

183 (182) Face largely honey-yellow with small brown or darker mark just below each antennal socket ( Fig. 31B ); codon at positions 1597–99 coding for N; codon at positions 1783–85 coding for I; codon at positions 1882–84 coding for N........ 184

- Face largely brown with clypeus (and sometimes small central area above clypeus), malar region, and narrow line next to eye ivory-white to yellowish ( Fig. 137B ); codon at positions 1597–99 coding for S; codon at positions 1783–85 coding for M; codon at positions 1882–84 coding for A............................................... A. prillae sp. nov.

184 (183) Lateral lobes of mesoscutum dark, middle lobe of mesoscutum with a brown to black mark at midlength ( Fig. 31F ); propodeum more coarsely sculptured; base 1869 C.................................... A. brunniguttatus sp. nov.

- Lateral lobes of mesoscutum variable, but middle lobe without brown transverse mark at midlength ( Fig. 45C ); propodeum more finely sculptured; base 1869 T................................................................. 185

185 (184) Mesoscutum largely honey coloured except narrowly brown-black posterolaterally ( Fig. 45C ); metasomal tergite 2 with a large cream coloured mark medio-posteriorly, often extending to postero-lateral part of tergite and sometimes to midanterior of tergite ( Fig. 45F ); bases 1557, 1830, 1875 and 2079 T, G, C and A respectively....... A. complexus sp. nov.

- Mesoscutum more extnsively dark marked, least with lateral lobes largely brown to black; metasomal tergite 2 with only a small rufous mark medio-posteriorly ( Fig. 84F ); bases 1557, 1830, 1875 and 2079 C, A, A and G respectively............................................................................................ A. herrena sp. nov.

186 (182) Metasoma entirely black ( Fig. 69F ); mesoscutum black except for midposterior part of middle lobe ( Fig. 69C )............................................................................................. A. faciei sp. nov.

- Coloration not as above.......................................................................... 187

187 (186) Third segment of maxillary palp swollen, only 4 x longer than wide ( Fig. 183B ); side of mesopleuron largely evenly foveate-rugose ( Fig. 183E )....................................................... A. turgidipalpus sp. nov.

- Third segment of maxillary palp slender ( Figs 179B ); side of mesopleuron usually aciculate with foveate or rugose sculpture restricted to dorsal parts and precoxal sulcus.................................................. 188

188 (187) Basolateral flange of 1st metasomal tergite not strongly and angularly produced and not concave posteriorly, forming a smooth lobe virtually reaching spiracle ( Figs 154E,G ) AND antenna with more than 40 flagellomeres............. 189

- Baso-lateral flange of 1st metasomal tergite angularly produced and distinctly concave posteriorly ( Fig. 36F ) OR antenna with fewer than 39 flagellomeres................................................................... 192

189 (188) Hind wing vein 2-SC+R strongly transverse ( Fig. 179E ).................................. A. transvena sp. nov.

- Hind wing vein 2-SC+R not transverse.............................................................. 190

190 (189) Metasoma almost entirely dark brown to black except for paler brown to cream coloured area at the base of tergite 2 ( Fig. 154G ); propodeum with very coarse rugose sculpture ( Fig. 154E ) and with distinct, rather ridge-like posterolateral protuberances ( Fig. 154D ); metasomal tergite 3 very shiny with longitudinal striation almost parallel ( Fig. 154G ).......................................................................................... A. rivulus sp. nov.

- Metasoma with far more extensive pale markings ( Figs 9E , 99G ); propodeum less coarsely sculptured ( Fig. 99E ) and without distinct posterolateral ridge-like protuberances ( Fig. 99C ); longitudinal striation of tergite 3 usually distinctly curved and diverging from midline ( Figs 9E , 99G )..................................................... 191

191 (190) Antenna with fewer than 31 (29) flagellomeres; 3 rd metasomal tergite almost entirely black ( Fig. 99G )................................................................................................. A. magratae sp. nov.

- Antenna with more than 32 (34) flagellomeres; 3 rd metasomal tergite almost entirely white, narrowly black laterally ( Fig. 9E ).............................................................................. A. achingae sp. nov.

192 (188) 1 st metasomal tergite with inverted wine-glass shaped black mark.............................. A. calicis sp. nov.

- 1 st metasomal tergite not patterned as above............................................................ 193

193 (192) Propodeum entirely black ( Fig. 174D ).................................................. A. teatimei sp. nov.

- Propodeum pale yellow medially, black laterally ( Fig. 161F ).............................................. 194

194 (193) Frons virtually flat, without distinct depressions behind antennal sockets and without sublateral carina paralleling eye ( Fig. 161C,D )..................................................................... A. sesquipunctatus sp. nov.

- Frons distinctly depressed behind antennal sockets ( Figs 27D , 64C , 92D ) and with sublateral carina paralleling eye and sometimes then curving to form posterolateral border of frontal depressions.................................. 195

195 (194) Mesopleuron including precoxal sulcus more or less evenly aciculate, precoxal sulcus virually absent, indicated by only a weak depression with very faint striae or a very short, narrow line of just 3 or 4 small foveae, area below wings not or hardly longitudinally striate ( Figs 20C , 27C ).......................................................... 196

- Anterior of mesopleuron and precoxal sulcus conspicuousley more coarsely sculptured rugulose to striate, area of mesopleuron below wings longitudinally striate ( Figs 70D , 92C )........................................... 197

196 (195) Episternal scrobe distinct, area above it slightly swollen and specular area with distinctly reduced sculpure ( Fig. 27C ); codon at 1894–96 ATT coding for I; codon at 1972–74 TTA coding for L........................ A. binkyi sp. nov.

- Episternal scrobe virtually absent, area above it not swollen and specular area without differentiated sculpture ( Fig. 20C ); codon at 1894–96 GTT coding for V; codon at 1972–74 TTT coding for F...................... A. ascrobi sp. nov.

197 (195) Hind wing vein m-cu interstitial........................................................ A. liessa sp. nov.

- Hind wing vein m-cu antefurcal ( Fig. 64D )............................................................ 198

198 (197) Hind coxa with trace of fine oblique striation on basodorsal and basomedial surface; base 1901 G. A. divergerus sp. nov.

- Hind coxa without a trace of oblique striation on basodorsal and basomedial surface ( Fig. 70D ); base 1901 A........................................................................................ A. flannelfooti sp. nov.

199 (169) Hind wing without vein m-cu ( Fig. 99F )............................................................... 00

- Hind wing with vein m-cu (see Fig. 154F )............................................................. 205

200 (199) Antenna with fewer than 31 (29) flagellomeres.......................................... A. magratae sp. nov.

- Antenna with more than 31 (33–40; precise number unknown in rincewindi sp. nov.) flagellomeres............... 201 201 (200) Propodeum entirely black ( Figs 63C , 152D , 167E ); codon at 1807–09 coding for V or M; codon at 1825–27 coding for S or V............................................................................................. 202

- Propodeum largely pale, at least broadly anteriorly and almost reaching posterior ( Figs 135D , 160A ); codon at 1807–09 coding for I; codon at 1825–27 coding for T.......................................................... 204

202 (201) Face with large brown mark, occipital carina dorso-laterally brown to dark brown; codon at 1684–86 ciding for I; codon at 1714–16 coding for V; codon at 1789–91 coding for A; codon at 1825–27 coding for S............ A. stohelit sp. nov.

- Face and back of head uniformly pale yellow; codon at 1684–86 ciding for I; codon at 1714–16 coding for V; codon at 1789–91 coding for S; codon at 1825–27 coding for V................................................... 203

203 (202) Codon at 1798–1800 coding for L; codon at 1807–09 coding for M; codon at 310–312 coding for V.................................................................................................. A. dimorphus sp. nov.

- Codon at 1798–1800 coding for F; codon at 1807–09coding for V; codon at 310–312 coding for I. A. rincewindi sp. nov.

204 (201) Antenna with more than 37 [39] flagellomeres; fore wing vein 2-CU1 1.3 x 1-CU1 ( Fig. 160E ); codon at 1657–59 coding for I; codon at 2044–46 coding for I................................................... A. selachiii sp. nov.

- Antenna with fewer than 37 [35] flagellomeres; fore wing vein 2-CU1 1.2 x 1-CU1 ( Fig. 135E ); codon at 1657–59 coding for V; codon at 2044–46 coding for V................................................... A. ponderi sp. nov.

205 (199) 3 rd metasomal tergite of female with broad transverse black band that is produced into a point medioposteriorly ( Figs 39F , 85F , 94F )..................................................................................... 206

- 3 rd metasomal tergite of female variably coloured but not as above......................................... 208

206 (205) Codon at 1816–1818 coding for V.................................................. A. lipwigi sp. nov. (part)

- Codon at 1816–1818 coding for I................................................................... 207

207 (206) 1 st metasomal tergite with a pale median line ( Fig. 39F ); codon at 1825–27 coding for T; codon at 2041–43 GCA coding for A............................................................................... A. caprinus sp. nov.

- 1 st metasomal tergite solid dark brown-black on basal 0.8 ( Fig. 85F ); codon at 1825–27 coding for M; codon at 2041–43 ACA coding for T..................................................................... A. hircus sp. nov.

208 (205) Basal lobes of 1 st metasomal tergite more or less evenly rounded, posteriorly not or hardly emarginate, or if weakly emarginate, more or less straight anteriorly, the steepest angles (i.e. most close to transverse) converging posteriorly at 20–45 degrees ( Figs 94E , 146F , 166F )................................................................ 209

- Basal lobes of 1 st metasomal tergite more strongly and sharply protruding, posteriorly strongly emarginate, the steepest angles (i.e. most close to transverse) converging posteriorly at more than 90 degrees ( Figs 89E , 109E )............. 214

209 (208) Body entirely or almost entirely cream to yellow, at most with a few brown patches on propodeum and 1 st and 2 nd metasomal tergites ( Figs 151E , 172E )................................................................ 210

- Body boldly marked with dark brown or black, tergites 2 and 3 at least with large black lateral stripes ( Figs 93E , 94F ) 211

210 (209) Antenna with fewer than 39 [37–38] flagellomeres; fore wing vein 3-SR less than 1.5 [1.25] x vein r ( Fig. 172A ); hind wing vein 2-SC+R longitudinal...................................................... A. sutthisani sp. nov.

- Antenna with more than 39 [40] flagellomeres; fore wing vein 3-SR more than 1.5 [1.8] x vein r ( Fig. 151A ); hind wing vein 2-SC+R virtually interstitial...................................................... A. ridcullyi sp. nov.

211 (209) Fore wing 2 nd submarginal cell very elongate, vein 3-SR more than 2 (2.3) x length of 2-SR ( Fig. 166E ); middle lobe of mesoscutum with a pair of longitudinal brown lines ( Fig. 166F ).............................. A. stibbonsi sp. nov.

- Fore wing 2 nd submarginal cell not elongate, vein 3-SR less than than 2 (1.5–1.7) x length of 2-SR ( Fig. 146E ); middle lobe of mesoscutum either uniformly pale yellow ( Fig. 146D ) or with a single brown patch ( Fig. 94D )................. 212

212 (211) Pterostigma largely dark grey brown, pale cream brown basally and apically ( Fig. 94A ); fore wing vein 3-SR 1.7 x 2-SR; fore wing veins 2-SR and 2-M forming an angle of approximately 75 degrees................................. 213

- Pterostigma with basal 0.4 pale, distal part dark grey brown ( Fig. 146E ); fore wing vein 3-SR 1.75 x 2-SR ( Fig. 146E ); fore wing veins 2-SR and 2-M forming an angle of approximately 85 degrees ( Fig. 146E )......... A. pteppicymoni sp. nov.

213 (212) Codon at 1792–4 ATA coding for M..................................................... A. lipwigi sp. nov.

- Codon at 1792–4 ACA coding for T............................................... A. lipwigduplicitus sp. nov.

214 (208) 2 nd metasomal tergite approximately 1.35 x wider posteriorly than long ( Fig. 89E ); codon at 1594–96 ATA coding for M; codon at 1606–08 GTT coding for V; codon at 1714–16 GTT coding for V................... A. klopfsteinae sp. nov.

- 2 nd metasomal tergite less than 1.2 (1.15 or less) x wider posteriorly than long ( Figs 51C , 158F ); codon at 1594–96 ACA coding for T; codon at 1606–08 ATT or ATC coding for I; codon at 1714–16 ATT coding for I.................. 215

215 (214) Ocelli rather large, distance across posterior ocelli approximately 4 x shortest distance between posterior ocellus and eye................................................................................................ 216

- Ocelli rather smaller, distance across posterior ocelli approximately 2.5–3.0 x shortest distance between posterior ocellus and eye ( Figs 62B , 101D )......................................................................... 218

216 (215) 3 rd tergite coarsely striate ( Fig. 51B ); codon at position 1930–32 ACA coding for T; codon at 1972–74 TTA coding for L.............................................................................. ... A. contemptus sp. nov.

- 3 rd tergite finely striate ( Figs 158F ); codon at position 1930–32 GCA coding for A; codon at 1972–74 TTC or TTT coding for F......................................................................................... 217

217 (216) Propodeum densely setose with fine rugulose sculpture ( Fig. 158D ); codon at 1720–22 GTA coding for V; codon at position 1807–1809 ATT coding for I................................................. A. sacharissa sp. nov.

- Propodeum less densely setose, with coarser sculpture ( Fig. 109D ); codon at 1720–22 ATT coding for I; codon at position 1807–1809 ACT coding for T............................................................ A. morti sp. nov.

218 (215) Metapleuron cream ( Fig. 62A ); codon at 1972–74 TTA coding for L........................... A. deyoyoi sp. nov.

- Metapleuron brown ( Fig. 101C ); codon at 1972–74 TTC coding for F........................... A. malichi sp. nov.

Figures

FIGURE 2. Scanning electron micrographs illustrating key features. A–D, showing forms of hind claw pecten, A, A. coronarius group unidentified species (voucher BCLDQ01248); B, A. antefurcalis sp. nov.; C, A. pectopulicis sp. nov. (voucher BCLDQ00211); D, A. calvus sp. nov. (voucher BCLDQ01483); E–F, showing absence and presence of comb of specialised adpressed setae apicomedially and apicoventrally on hind tibia, E, Aleiodes nivori sp. nov. (voucher BB0005); F, A. calvus sp. nov. (voucher BCLDQ01483).

FIGURE 3. Scanning electron micrographs illustrating key features. A, inner aspect of apex of hind tibia and hind basitarsus of Yelicones sp.; B, A. sp., dorsal view of hind coxa (voucher BCLDQ0295); C, A. parisaae sp. nov., dorsal view of hind coxa; D, Heterogamus sp., dorsal view of hind coxa (voucher BCLDQ0194).

FIGURE 4. Scanning electron micrographs illustrating key features. A, elongate penultimate flagellomes of A. sesquipunctatus sp. nov. (voucher BCLDQ, 1481); B, penultimate flagellomeres of A. brunniguttatus sp. nov. (voucher BCLDQ00678); C, ventral area of median flagellomeres of A. pronopus sp. nov. (voucher BCLDQ00195); ventral area of median flagellomeres of Heterogamus sp. showing broad absence of longitudinal placode sensilla.

FIGURE 9. A. achingae sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma; F, fore wing.

FIGURE 10. A. aciculodeum sp. nov. A, habitus; B,face, C, head and anterior mesosoma, dorsal view; D, mesosoma, lateral view; E, propodeum and 1st tergite, dorsal view; F, metasoma.

FIGURE 11. A. adorabelleae sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 12. A. agagg sp. nov. A, habitus; B,face, C, head and anterior mesosoma, lateral view; D, metasoma; E, head and anterior mesosoma, dorsal view.

FIGURE 13. A. alboluteus sp. nov. A, habitus; B, face; C, head, lateral view; D, head and mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 14. A. anguaae sp. nov. A, habitus; B, face; C, head, lateral view; D, head and mesosoma, dorsal view; E, mesosoma, lateral view; F, propodeum and 1st tergite, dorsal view; G, metasoma.

FIGURE 15. A. angulodeum sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D, head, dorsal view; E, propodeum; F, metasoma.

FIGURE 16. A. antescutum sp. nov. A, habitus; B, head and anterior mesosoma, dorsal view; C, paratype with associated mummified host caterpillar remains; D, metasoma; E, head and mesosoma, lateral view.

FIGURE 18. A. archicolorus sp. nov. A, habitus; B, face; C, head, dorsal view; D, anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, propodeum and 1st tergite, dorsal view; G, metasoma

FIGURE 19. A. areelucki sp. nov. A, habitus; B, head, lateral view; C, mesosoma, lateral view; D, head and mesosoma, dorsal view; E, metasoma.

FIGURE 20. A. ascrobi sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head, dorsal view; E, anterior mesosoma, dorsal view; F, propodeum, dorsal view; G, metasoma.

FIGURE 21. A. asperum sp. nov. A, habitus; B, face; C, head, lateral view; D, head and mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma, oblique view.

FIGURE 22. A. atuin sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, fore wing; E, metasoma; F, metanotum and propodeum, dorsal view.

FIGURE 23. A. bachmaduplus sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D, head, lateral view; E, G, mesosoma lateral view, holotype and paratype respectively; F, H, metasoma, holotype and paratype respectively.

FIGURE 24. A. bachmatriplus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, lateral view; E, metasoma; F, head and anterior mesosoma, dorsal view.

FIGURE 25. A. basipunctatus sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D mesosoma, lateral view; E, metasoma.

FIGURE 26. A. basistriatus sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 27. A. binkyi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 28. A. biscutus sp. nov. A, habitus; B, face; C, head, lateral view; D, mesosoma, lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 29. A. bobwhartoni sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, wings; E, apex of hind tibia, ventral view; F, metasoma.

FIGURE 30. A. brevimedius sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, metasoma, dorsal view; F, metasoma, lateral view; G wings, H, metasoma.

FIGURE 31. A. brunniguttatus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, lateral view; E, head, dorsal view; F, anterior mesoscutum, dorsal view; G, fore wing; H, metasoma.

FIGURE 32. A. butcheri sp. nov. A, habitus; B, face; C, head, dorsal view; D, head, lateral view; E, mesosoma, dorsal view; F, wings; G, metasoma, dorsal view; H, metasoma, lateral view.

FIGURE 33. A. buzuriduplus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 34. A. buzuriquadruplus sp. nov. A, habitus; B, wings; C, mesosoma, dorsal view; D, metasoma.

FIGURE 35. A. buzuritriplus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 36. A. calicis sp. nov. A, habitus; B, face; C, mesosoma, dorsal view; D, head, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 37. A. calvus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, metasoma; F, mid posterior part of fore wing showing glabrous zone below vein 1-CU1; G, basal of hind wing showing glabrous subbasal cell; H, anterior mesosoma, dorsal view;

FIGURE 38. A. canus sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 39. A. caprinus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 40. A. castaneus sp. nov. A, female habitus; B, male habitus; C, face; D, head and mesosoma, lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 41. A.centralis sp. nov. A, habitus; B, head, dorsal view; C, mesoscutum and scutellum, dorsal view; D, hind coxa, dorsal view; E, metasoma.

FIGURE 42. A. chamba sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, head, lateral view; E, mesosoma, lateral view; F, fore wing; G, metasoma.

FIGURE 43. A–C, Aleiodes chenduplus sp. nov.; D–F, A. cheni sp. nov. A, habitus; B, head and anterior mesosoma, dorsal view; C, metasoma; D, head, dorsal view; E, mesosoma, lateral view; F, propodeum and 1st tergite, dorsal view.

FIGURE 44. A. codon sp. nov. A, habitus; B, head and anterior mesosoma, dorsal view; C, head and mesosoma, lateral view; D, fore wing; E, metasoma.

FIGURE 45. A. complexus sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, head, lateral view; E, mesosoma, lateral view; F, metasoma.

FIGURE 46. A. concoronarius sp. nov. A, habitus; B, head, dorsal view; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 47. A. conina sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, lateral view; E, head and anterior mesosoma, dorsal view; F, propodeum and 1st tergite, dorsal view; G, metasoma.

FIGURE 48. A. connudatum sp. nov. A, habitus; B, head, dorsal view; C, head and mesosoma, lateral view; D, mesosoma, dorsal view; E, fore wing; F, base of hind wing showing glabrous subbasal cell; G, metasoma.

FIGURE 49. A. conpectenus sp. nov. A, habitus; B, head, lateral view; C, mesosoma, lateral view; D, head, dorsal view; E,

FIGURE 50. A. constriatum sp. nov. A, habitus; B, head, lateral view; C, mesosoma, lateral view; D, head and anterior mesosoma, oblique dorsal view; E, fore wing; F, metasoma, arrow indicating moderately strongly produced, subangular basal lobes of 1st tergite.

FIGURE 51. A. contemptus sp. nov. A, mesosoma, dorsal view; B, face; C, metasoma; D, head and mesosoma, lateral view; E, fore wing.

FIGURE 52. A. convexus. A, habitus; B, face; C, mesosoma, lateral view; D, head, lateral view; E, anterior mesosoma, dorsal view; F, metasoma.

FIGURE 54. A. corrusciput sp. nov. A, habitus; B, face; C, mesoscutum, dorsal view; D, head and pronotum dorsal view; E, metasoma, lateral view; F, metasoma.

FIGURE 55. A. cramum sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, anterior mesosoma, dorsal view; F, metasoma.

FIGURE 56. A. curtivena sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, oblique dorsal view; E, metasoma.

FIGURE 57. A. curvicauda sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, metasoma, lateral view; F, metasoma; G, propodeum.

FIGURE 58. A. damus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 59. A. darlingtonia sp. nov. A, habitus; B, head, dorsal view; C, mesosoma, lateral view; D, mesosoma, dorsal view; E, metasoma, dorsal view; F, metasoma, lateral view.

FIGURE 60. A. deathi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 61. A. definus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and pronotum, lateral view; E, head and pronotum, dorsal view; F, metasoma.

FIGURE 62. A. deyoyoi sp. nov. A, habitus; B, head, dorsal view; C, head and mesosoma, lateral view; D, anterior mesosoma, dorsal view; E, metasoma.

FIGURE 63. A. dimorphus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and pronotum, lateral view; E, anterior mesosoma, dorsal view; F, metasoma.

FIGURE 64. A. divergerus sp. nov. A, habitus; B, face; C, head, dorsal view; D, wings; E, mesosoma, dorsal view, arrow indicating medium-ized, posteriorly steeply concave basal lobe of 1st tergite; F, metasoma.

FIGURE 65. A. downeyi sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, wings; F, propodeum and 1st tergite, dorsal view; G, metasoma.

FIGURE 66. A. eadyi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 67. A. elongatus sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D, head, lateral view; E, mesosoma, latteral view; F, metasoma.

FIGURE 68. A. etvalinus sp. nov. A, habitus; B, face, C, head and anterior mesosoma, dorsal view; D, head, lateral view; E, mesosoma, latteral view; F, wings; G, metasoma.

FIGURE 69. A. faciei sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 70. A. flannelfooti sp. nov. A, habitus; B, face; C, head, dorsal view; D, head and mesosoma, lateral view; E, mesosoma, dorsal view; F, metasoma.

FIGURE 71. A. flavostriatus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and mesosoma, dorsal view; E, metasoma.

FIGURE 72. A. fovodeum sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, mesosoma, dorsal view; E, wings; F, metasoma.

FIGURE 73. A. fuscomedius sp. nov. A, habitus; B, face, C, head and anterior mesosoma, dorsal view; D, hea, lateral view;

FIGURE 74. A. gaga sp. nov. A, face; B, mesosoma, lateral view; C, habitus; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 75. A. gaspodei sp. nov. A, habitus; B, face, oblique view; C, mesosoma, lateral view; D, head and pronotum, dorsal view; E, fore wing; F, mesosoma, dorsal view; G, metasoma.

FIGURE 76. A. georgiaae sp. nov. A, habitus; B, face; C, head, lateral view; D, mesosoma, lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 77. A. glabribasalis sp. nov. A, habitus; B, head, dorsal view;; C, mesosoma, lateral view; D, mesosoma, dorsal view; E, fore ing; F, metasoma.

FIGURE 78. A.globifemurus sp. nov. A, habitus; B, face; C, head, lateral view; D, anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 79. A. glutinum sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, anterior mesosoma, dorsal view; F, metasoma.

FIGURE 80. A. griseimaculus sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, metasoma.

FIGURE 81. A. guidaae sp.nov. A, habitus;B, face; C, mesosoma,lateral view; D, head and anterior mesosoma,dorsal view; E, metasoma.

FIGURE 82. A. hei sp. nov. A, habitus; B, face; C, head and mesoscutum, dorsal view; D, head, lateral view; E, mesosoma, lateral view; F, propodeum, dorsal view; G, wings; H, metasomal tergites 2 and 3.

FIGURE 83. A. helenhippersonsae sp. nov. A, head and mesoscutum, lateral view; B, metasoma, lateral view; C, propodeum and 1st tergite, dorsal view; D, metasoma.

FIGURE 84. A. herrena sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 85. A. hircus sp. nov. A, mesosomal, lateral view; B, face; C, head and mesoscutum, dorsal view; D, scutellum and propodeum, dorsal view; E, wings; F, metasoma.

FIGURE 86. A. incisus sp. nov. A, habitus; B, face; C, head and mesoscutum, dorsal view; D, mesosoma, lateral view; E, anterior mesosoma, dorsal view; F, propodeum and 1st tergite, dorsal view; G, metasoma.

FIGURE 87. A. jimwhitfieldi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, metasoma.

FIGURE 88. A. kitchingi sp. nov. A, habitus; B, mesosoma, lateral view; C, head, lateral view; D, head, dorsal view; E, wings; F, metasoma.

FIGURE 89. A. klopfsteinae sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, head and mesosoma, lateral view; E, metasoma.

FIGURE 90. A. lavaeolus sp. nov. A, habitus; B, head, dorsal view; C, head and mesosoma, oblique lateral view; D, mesosoma, dorsal view; E, metasoma.

FIGURE 91. A. lemniscus sp. nov. A, habitus; B, face; C, head, lateral view; D, head and mesoscutum, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 92. A. liessa sp. nov. A, habitus; B, head and mesosoma, dorsal view; C, mesosoma, lateral view; D, head, lateral view; E, metasoma.

FIGURE 93. A. lipwigduplicitus sp. nov. A, habitus; B, head, dorsal view; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 94. A. lipwigi sp. nov. A, habitus; B, head, dorsal view; C, head and mesosoma, lateral view; D, anterior mesosoma, dorsal view; E, propodeum, dorsal view; F, metasoma.

FIGURE 95. A. lobocarinus sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D, head and mesosoma, lateral view; E, metasoma.

FIGURE 96. A. longivena sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, propodeum and 1st tergite, dorsal view; E, wings; F, metasoma.

FIGURE 97. A. macroophthalmos sp. nov. A, habitus; B, head, dorsal view; C, mesosoma, lateral view; D, head, lateral view; E, propodeum and 1st tergite, dorsal view; F, metasoma; G, wings.

FIGURE 98. A. maculiput sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D, metasoma.

FIGURE 99. A. magratae sp. nov. A, habitus; B, head, dorsal view; C, mesosoma, lateral view; D, anterior mesosoma, dorsal view; E, propodeum and 1st tergite, dorsal view; F, hind wing showing absence of vein m-cu; G, metasoma.

FIGURE 100. A. malarius sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D, metasoma.

FIGURE 101. A. malichi sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 103. A. mediomaculatus sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, fore wing; G, metasoma.

FIGURE 104. A. megaophthalmos sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, anterior mesosoma, dorsal view; F, mesosoma, lateral view; G, metasoma.

FIGURE 105. A. mellificus sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, mesosoma, lateral view; F, anterior mesosoma, dorsal view; G, metasoma.

FIGURE 106. A. mericeti sp. nov. A, habitus; B, head and pronotum, dorsal view; C, mesosoma, dorsal view; D, fore wing; E, head and mesosoma, lateral view; F, metasoma.

FIGURE 107. A. microophthalmos sp. nov. A, habitus; B, face; C, head and mesosoma, dorsal view; D, metasoma.

FIGURE 108. A. molecryptus sp. nov. A, habitus; B, face; C, head, dorsal view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; E, metasoma.

FIGURE 109. A. morti sp. nov. A, habitus; B, face; C, wings, D, mesosoma, dorsal view; E, metasoma.

FIGURE 110. A. mutilus sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 111. A. narangae. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, head and anterior mesosoma, lateral view; E, metasoma; F, details mor metasomal tergites 2 and 3.

FIGURE 112. A. nathismus sp. nov. A, habitus; B, head, lateral view; C, head and anterior mesosoma, dorsal view; D, mesosoma lateral view; E, head, dorsal view; F, metasoma.

FIGURE 113. A. necsubson sp. nov. A, habitus; B, face; C, head, lateral view; D, mesosoma lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 114. A. niveni sp. nov. A, habitus; B, face; C, mesosoma lateral view; D, head and anterior mesosoma, dorsal view; E, apical part of hind tarsus, arrow heads indicating positions of robust, yellow pecten teeth; F, metasoma.

FIGURE 115. A. nivori sp. nov. A, habitus; B, face; C, mesosoma lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 116. A. nonicones sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, fore wing; E, propodeum and 1st tergite, dorsal view; F, metasoma.

FIGURE 117. A. occimaculatus sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, mesosoma lateral view; E, metasoma.

FIGURE 118. A. opus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, propodeum; F, anterior mesosoma, dorsal view; G, metasoma.

FIGURE 119. A. orion sp. nov. A, habitus; B, mesosoma, lateral view; C, head and anterior mesosoma, dorsal view; D, metasoma.

FIGURE 120. A. paenicarinus sp. nov. A, habitus; B, head, dorsal view; C, mesosoma, lateral view; D, head, lateral view; E, metasoma.

FIGURE 121. A. pallimedius sp. nov. A, habitus; B, head and anterior mesosoma, dorsal view; C, head and anterior mesosoma, lateral view; D, fore wing; E, metasoma and hind femur.

FIGURE 122. A. palmae sp. nov. A, habitus; B, face; C, head and pronotum, dorsal view; D, mesosoma, lateral view; E, mesoscutum; F, metasoma.

FIGURE 123. A. palmatipes sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, mesosoma, lateral view; F, mesoscutum, oblique view; G, metasoma.

FIGURE 124. A. parabuzurae sp. nov. A, habitus; B, face; C, head, lateral view; D, mesosoma, lateral view; E, metasoma.

FIGURE 125. A. parasongsi sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, mesosoma, lateral view; F, mesosoma, dorsal view [note: right half and posterior of propodeum obscured by glue]; G, metasoma.

FIGURE 126. A. parisaae sp. nov. A, habitus; B, face; C, head, lateral view; D, mesosoma, lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma, lateral view; G,, metasoma, dorsal view.

FIGURE 127. A. paulmashi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, oblique dorsal view; E, fore wing; F, metasoma

FIGURE 128. A. pectopulicis sp. nov. A, habitus; B, face; C, head, oblique dorsal view; D, mesosoma, lateral view; E, mesosoma, ventral view; F, metasoma.

FIGURE 129. A–C, Aleiodes pectunguis sp. nov., D–G, A. pectunguisella sp. nov. A, habitus; B, head and anterior mesosoma, dorsal view; C, propodeum and anterior metasoma, dorsal view; D, habitus, E, face; F, anterior mesosoma, dorsal view; G, metasoma.

FIGURE 130. A. penultimoluteus sp. nov. A, antenna of male; B, habitus; C, face; D, head, dorsal view; E, mesosoma, lateral view; F, head and anterior mesosoma, dorsal view; G, metasoma.

FIGURE 131. A. phantasmatis sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 132. A. pinnulae sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, F, metasomas.

FIGURE 133. A. placidus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, wings with arrows indicating relatively short vein 1-M relative to M+CU of hind wing; F, metasoma.

FIGURE 134. A. polititergus sp. nov. A, habitus; B, head and anterior mesosoma, dorsal view; C, head and pronotum, lateral view; D, anterior mesosoma, dorsal view; E, mesosoma lateral view; F, propodeum and anterior of 1st tergite, dorsal view; G, metasoma.

FIGURE 135. A. ponderi sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, propodeum and anterior of 1st tergite, dorsal view; E, wings; F, metasoma.

FIGURE 136. A. postmaculus sp. nov. A, habitus; B, face; C, head, dorsal view; D, mesosoma lateral view; E, mesosoma, dorsal view; F, metasoma.

FIGURE 137. A. prillae sp. nov. A, habitus; B, face; C, head and pronotum, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma; F, mesosoma lateral view.

FIGURE 138. A. probuzurae sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, mesosoma lateral view; E, metasoma.

FIGURE 140. A. procoronarius sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, head dorsal view; E, mesosoma lateral view; F, metasoma.

FIGURE 141. A. pronopus sp. nov. A, habitus; B, face; C, head, dorsal view; D, anterior mesosoma, dorsal view; E, metasoma.

FIGURE 143. A. propodocarinus sp. nov. A, habitus; B, face; C, head and anterior mesosoma, lateral view; D, head dorsal view; E, mesosoma and 1st tergite, dorsal view; F, metasoma.

FIGURE 144. A. protocastaneus sp. nov. A, habitus; B, face; C, head, lateral view; D, head dorsal view; E, metasoma; F, mesosoma lateral view.

FIGURE 145. A. pseudicones sp. nov. A, habitus; B, face; C, fore leg; D, anterior mesosoma, dorsal view; E, mesosoma lateral view; F, metasoma.

FIGURE 146. A. pteppycomi sp. nov. A, habitus; B, face; C, head and anterior mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 147. A. ptraci sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 148. A. rectanguliguttatus sp. nov. A, habitus; B, upper part of head, lateral view; C, head and anterior mesosoma, dorsal view; D, mesosoma, lateral view; E, metasoma.

FIGURE 149. A. reticulisoma sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 150. A. reus sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 151. A. ridcullyi sp. nov. A, habitus; B, face; C, head and anterior mesosoma, dorsal view; D, mesosoma lateral view; E, metasoma.

FIGURE 152. A. rincewindi sp. nov. A, habitus; B, face, oblique view; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 153. A. risaae sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, hind coxa; F, metasoma, dorsal view; G, metasoma, lateral view.

FIGURE 154. A. rivulus sp. nov. A, habitus, holotype and paratype, upper and lower specimens respectively; B, face; C, head and anterior mesosoma, dorsal view; D, mesosoma, lateral view, propodeum, arrow indicating ridge-like posterolateral protuberance; E; F, hind wing showing presence of vein m-cu; G, metasoma.

FIGURE 155. A. rufomedius sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, wings.

FIGURE 157. A. rugoscutus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, mesosoma, dorsal view; F, head, lateral view; G, hind coxa; H, wings; I, metasoma.

FIGURE 158. A. sacharissa sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, mesosoma, dorsal view; E, wings; F, metasoma.

FIGURE 159. A. scottscawi sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 160. A. selachiii sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, anterior mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 161. A. sesquipunctatus sp. nov. A, habitus; B, face, oblique view; C, head, lateral view; D, head, oblique dorsal view; E, anterior mesosoma, dorsal view; F, propodeum; G, metasoma.

FIGURE 162. A. similicodon sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view, arrow indicating ‘V’- shaped carina on frons; E, mesosoma, lateral view; F, anterior mesosoma, dorsal view; G, metasoma.

FIGURE 163. A. songsi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, propodeum; F, wings; G, metasoma.

FIGURE 164. A. sophieae sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, lateral view; E, anterior mesosoma, dorsal view; F, metasoma.

FIGURE 165. A, B, Aleiodes spurivena; C–E, A. (Hemigyroneuron) roberti. A, habitus; B, fore wing; C, habitus D, hind wing; E, fore wing detail.

FIGURE 166. A. stibbonsi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, fore wing; F, mesosoma, dorsal view; G, metasoma and hind femora.

FIGURE 167. A. stohelit sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, head and anterior mesosoma, lateral view; F, metasoma.

FIGURE 168. A. subemarginatus sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 169. A. subfuscomedius sp. nov. A, habitus; B, face; C, head, lateral view; D, mesosoma, lateral view; E, head and anterior mesosoma, dorsal view; F, metasoma.

FIGURE 170. A. submediofuscus sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 171. A. subson sp. nov. A, male and female habitus, left and right respectively; B, face; C, mesosoma, lateral view; D, head, lateral view; E, metasoma.

FIGURE 172. A. sutthisani sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 173. A. synorion sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and pronotum, dorsal view; E, mesosoma, dorsal view; F, fore wing; G, metasoma.

FIGURE 174. A. teatimei sp. nov. A, habitus; B, face; C, metasoma, arrow indicating moderately developed, posteriorly concave basal lobe of 1st tergite; D, mesosoma, dorsal view; E, fore wing.

FIGURE 175. A. tetrarugulosus sp. nov. A, habitus; B, head, lateral view; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 176. A. thirakupti sp. nov. A, habitus; B, face; C, head, lateral view; D, head, dorsal view; E, mesosoma, lateral view; F, mesosoma, dorsal view; G, metasoma.

FIGURE 177. A. tmaliaae sp. nov. A, habitus; B, face; C, head and pronotum, lateral view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma, oblique view.

FIGURE 178. A. tobiasi sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, metasoma.

FIGURE 179. A. transvena sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesoscutum, dorsal view; E, hind wing; F, metasoma.

FIGURE 180. A. tricoloripes sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and anterior mesosoma, dorsal view; E, wings; F, metasoma; G, propodeum, oblique view.

FIGURE 181. A. trisphaeropyx sp. nov. A, habitus; B, head and anterior mesoscutum, dorsal view; C, mesosoma, lateral view; Dm hind coxa; E, metasoma, dorsal view; F, metasoma, lateral view.

FIGURE 182. A. tulipus sp. nov. A, habitus; B, head, dorsal view; C, head and pronotum, lateral view; D, mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 183. A. turgidipalpus sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 185. Showing the difference in the degree of posterolateral emargination of the 3rd metasomal tergite. A. A. valinus sp. nov., B, A. incisus sp. nov.

FIGURE 186. A. variifemorus sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head, dorsal view; E, metasoma; F, anterior mesosoma, dorsal view.

FIGURE 187. A. vettinerii sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, mesosoma, dorsal view; E, fore wing; F, metasoma.

FIGURE 188. A. vietuput sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and mesosoma, dorsal view; E, wings; F, metasoma.

FIGURE 189. A. yalaensis sp. nov. A, habitus; B, face; C, head, lateral view; D, head and anterior mesosoma, dorsal view; E, mesosoma, lateral view; F, metasoma.

FIGURE 190. A. zuburae sp. nov. A, habitus; B, face; C, mesosoma, lateral view; D, head and mesosoma, dorsal view; E, metasoma.

FIGURE 191. Aleiodes (Arcaleiodes) dresdeni sp. nov. A, habitus; B, face; C, head and mesosoma, lateral view; D, head and pronotum, dorsal view; E, metasoma

FIGURE 192. Aleiodes (Arcaleiodes) siamensis sp. nov. A, habitus; B, face; C, head and pronotum, lateral view; D, head and anterior mesosoma, dorsal view; E, propodeum and 1st metasomal tergite, dorsal view; F, metasoma.

FIGURE 193. Aleiodes (Arcaleiodes) vanachterbergi sp. nov. A, habitus; B, head, dorsal view; C, mesosoma, lateral view; D, anterior mesosoma, dorsal view; E, propodeum; F, metasoma.