Escaryus japonicus Attems, 1927 *

Dyachkov, Yurii V. & Bonato, Lucio, 2024, An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia, ZooKeys 1198, pp. 17-54 : 17

29. Escaryus japonicus Attems, 1927 *

Escaryus japonicus Attems 1927: 299.

Escaryus japonicus - Attems 1929: 96; Byzova and Chadaeva 1965: 333; Shinohara 1972: 66; Titova 1969: 165; 1972: 135; 1973: 113; Ghilarov and Perel 1973: 46; Molodova 1973: 67; Alekseeva 1974: 8; Kurcheva 1977: 45; Markelov and Mineeva 1981: 130; Ganin 1997: 105, 109, 112, 114, 121, 124, 126, 129, 134, 141; 2006: 501; Vorobiova 1999: 33; Farzalieva 2008: 67; Volkova 2016: 676; Dyachkov 2017: 454; Nefediev et al. 2017a: 11; 2017c: 222; Dyachkov and Tuf 2018: 296; Nefediev 2019: 25; Dyachkov and Farzalieva 2023: 229.

Type locality.

Japan: Hokkaido: “Todohokhe” ( Attems 1927).

Type series.

Syntypes: 2 specimens, including a male and a juvenile. Deposited in NHMW ( Ilie et al. 2009).

Diagnosis.

An Escaryus species with body length reaching ≥ 4.3 cm; clypeus with large plagulae; labral arc relatively shallow, with denticles long and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur with a small distal denticle, all other articles with very small bulges; 43-55 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment rectangular, ~ 2 × as long as wide; coxal pores of similar size, on both ventral and lateral sides of coxopleura; anal pores present.

Distribution.

Western Siberia: Sverdlovsk, Chelyabinsk, Tomsk, and Kemerovo oblasts, republics of Altai and Khakassia ( Byzova and Chadaeva 1965; Titova 1972, 1973; Farzalieva 2008; Dyachkov 2017; Nefediev et al. 2017a, 2017c; Nefediev 2019). Eastern Siberia: Republic of Buryatia and Krasnoyarsk krai ( Alekseeva 1974; Titova 1973; Vorobiova 1999), Magadan oblast ( Berman and Leirikh 2019). Far East: Amur oblast, Maritime and Khabarovsk krais, Sakhalin oblast (Sakhalin Isl.) ( Titova 1973; Ghilarov and Perel 1973; Molodova 1973; Kurcheva 1977; Markelov and Mineeva 1981; Ganin 1997, 2006). Outside Asian Russia: European Russia ( Titova 1973; Farzalieva 2008; Volkova 2016), eastern Kazakhstan ( Dyachkov and Tuf 2018), Mongolia ( Dyachkov and Farzalieva 2023), Northern China ( Takakuwa and Takashima 1949), and Japan ( Attems 1927).

Remarks.

The record from the Krasnoyarsk krai by Vorobiova (1999) was questioned by Nefediev et al. (2017a).