Scaphisoma tricoloratum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1435-1436

5.2.21.4. Scaphisoma tricoloratum nov.sp. ( Figs 238, 239 )

T y p e m a t e r i a l: Holotype 6: Philippines Baracatan, 1500 m Mindanao June, 27-29, 1977 M. Satô leg (MHNG). Paratypes: Mindanao, with the same data as the holotype, 9 66 (MHNG); Mainit Hot Springs, 28.VII.1970, leg. M. Satô, 2 66 (MHNG); Mindanao, Bukidnon Prov., 5 km N. of Malaybalay, 9 0 0 m, 12.V.1996, leg. Bolm, 4 66 (SMNS, MHNG, BZLA); Mindanao, 30 km E. of Malaybalay, 5-9.V.1996, leg. Bolm, 2 66 (SMNS, MHNG).

D e s c r i p t i o n: Length 2.20-2.50 mm, width 1.50-1.72 mm. Head ochreous. Pronotum ochreous, with basal margin narrowly darkened and its lobe more broadly widened, darkened area usually not extended to middle third of disc. Hypomera ochreous. Elytra reddish-brown to blackish on prevailing surface, elytra with or without lighter subhumeral areas. Apical fifth of elytra yellowish. Meso- and metaventrites with anepisterna and epimera brown, abdominal sternite 1 somewhat lighter, apical abdominal segments ochreous. Appendages ochreous. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 12/12: IV 55/9: V 80/9: VI 67/10: VII 72/10: VIII 63/8: IX 68/10: X 68/14: XI 85/15. Pronotum very finely and sparsely punctate, with lateral margins evenly rounded, lateral margin carinae hardly exposed in dorsal view, lateral striae with hardly visible punctures. Tip of scutellum well visible. Elytra with lateral margin carinae entirely and well visible in dorsal view, apical margins rounded, inner apical angles not prominent, situated posterior level of outer angles; sutural margin not raised, sutural striae shallow, starting posterior level of scutellar tip, weakly converging apically, very finely punctate; adsutural areas flat, densely and very finely punctate, with punctures smaller than those on elytral disc, lateral striae punctate. Elytra with fairly coarse discal punctation, punctures rather well delimited, punctures intervals mostly as large to twice as large as puncture diameters. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera shorter than intervals to mesocoxae, about four times as long as wide. Median part of metaventrite slightly convex, without impressions, flattened between metacoxae, with strigulate microsculpture, punctation evanescent. Lateral parts of metaventrite microsculptured, with hardly visible punctation; antecoxal puncture rows indistinct. Submesocoxal areas 0.04-0.05 mm, about as sixth of intervals to metacoxae, submesocoxal lines slightly convex, distinctly punctate. Metanepisternum flat, weakly narrowed anteriad, inner margin straight, suture shallow. Protibiae straight, mesotibiae distinctly curved, metatibiae hardly curved, narrower than mesotibiae.

M a l e: Protarsomeres 1 to 3 and mesotarsomeres 1 and 2 strongly widened, tarsomeres 1 about as wide as or wider than apices of tibiae. Abdomen with strigulate microsculpture, almost impunctate. Sternite 1 with hardly visible punctures on narrow basomedian area; submetacoxal areas 0.07 mm, about as third of intervals to apical margin of sternite 1, submetacoxal lines convex, fairly coarsely punctate. Sternite 6 impressed in middle. Aedeagus ( Fig. 238, 239 ) 1.46-1.95 mm long, asymmetrical. Basal bulb very large and moderately sclerotized, with ventral ridge. Apical process of median lobe with long, weakly inflexed and abruptly bent ventral branch, latter appearing in lateral view truncate at apex. Dorsal branch of apical process short, strongly narrowed, reaching midlength of ventral branch. Parameres with robust ventral apophysis and large, overlapping dorsal lobes. Internal sac complex, with four elongate, robust sclerites in basal part followed by dense rows of spines.

E t y m o l o g y: The species epithet is a Latin adjective, meaning three-coloured.

C o m p a r a t i v e n o t e s: This species shares with S. tricolor most external diagnostic characters. Fully coloured males of S. tricoloratum differ by the darkened pronotal lobe. In addition, unlike to S. tricolor, the elytral surface anterior light apical areas may be blackish in this species. The shape of the large ventral branch of the apical process of the median lobe is quite different from that of other members of the group.

Figures

Figs 238 - 241: Aedeagi of Scaphisoma in dorsal and lateral views; (238, 239), S. tricoloratum nov. sp., in lateral view, without right paramere; (240, 241), S. tricolorinotum nov. sp. Scale = 0.3 mm.