Scaphisoma trimaculatum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1441-1442

5.2.21.10. Scaphisoma trimaculatum nov.sp. ( Figs 255 , 256 )

T y p e m a t e r i a l: Holotype 6: Luzon: Philippines Banga-an (1650m) Sagada Mountain Pv. 21.VII.1985 M. Sakai leg. (EUMJ). Paratypes: with the same data as the holotype, 2 66, 6♀♀ (EUMJ, MHNG); Luzon, env. Sagada St. Joseph Resthouse, 15-19.XII.1979, leg. L. Deharveng & J. Orousset, 1♀ (MHNG); Luzon, Mountain Prov., Sagada 1500 m, 22.VII.1985, leg. M. Sakai, 1♀ (EUMJ).

Description: Length 2.50-2.80 mm, width 1.72-1.90 mm. Head ochreous. Pronotum blackish with reddish or ochreous lateral areas. Hypomera ocreous. Elytra each blackish-brown to black along basal margin, on adsutural area, along sutural stria, along lateral margin, and with dark to blackish transverse band situated anterior light apical area; area between dark base and dark transverse band reddish; apical fourth of elytra yellowish. Meso- and metaventrite brown to blackish. Antennae and legs light brown to ochreous. Abdomen brown, with light apical segments. Length/width ratio of antennomeres as: III 18/11: IV 72/10: V 97/10: VI 83/10: VII 90/17: VIII 73/10: IX 87/14: X 78/14: XI 83/15. Pronotum fairly coarsely punctate on and nearby basal lobe. Upper part of hypomera with or without distinct punctures and pubescence. Elytra with sutural striae parallel in short anterior section, gradually converging from mid-third of sutural length to apices. Submesocoxal areas 0.05 mm, about as sixth of intervals to metacoxae. Sternite 1 all over extremely finely punctate, with submetacoxal areas 0.08 mmn almost as third of intervals to apical margin. Most of other external characters similar to those in S. tricoloratum, S. tridens, and S. tricoloripenne.

M a l e: Dark mesal area of pronotum restricted on to middle third of disc or forming narrow band, prevailing surface of pronotum ochreous or reddish. Apical margins of elytra truncate, oblique, distinctly crenulated, with inner angles situated posterior level of outer angles. Protarsomeres 1 strongly widened, about as wide as apices of protibiae, protarsomeres 2 somewhat narrower, protarsomeres 3 weakly widened. Mesotarsomeres 1 and 2 strongly widened, mesotarsomeres 1 distinctly wider than apices of mesotibiae. Sternite 6 flat in middle. Aedeagus ( Figs 255 , 256 ) 1.68 mm long. Apical process of median lobe with ventral branch long, inflexed and gradually narrowed apically, with tip curved and acute. Dorsal branch of median lobe short, hardly extended to level of midthird of ventral branch. Internal sac with large curved proximal sclerite, two moderately long admesal sclerites, central vesicle bearing fine denticles, and very dense rows of apical spine-like structures.

F e m a l e: Dark mesal areas of pronotum extended to cover most of discal surface, ochreous areas forming well delimited lateral spots. Apical margins of elytra strongly oblique and truncate, with inner angles denticulate, situated far posterior outer angles.

E t y m o l o g y: The species epithet is a Latin adjective, meaning three-spotted.

Comparative notes: The species resembles by its colour pattern to S. pulchrum, S. tricoloripenne and S. trifurcatum. It may be easily separated from S. pulchrum by the shortened sutural striae and by distinctly broader light areas of the elytra. The males of this new species differ drastically by the shape of the long apical process of the aedeagus. The females of S. pulchrum are unknown, those of S. tricoloripenne have strongly oblique apical margins of elytra, unlike females in S. trimaculatum. The females of S. trifurcatum have rounded inner apical angles of the elytra while the males of the latter species are unique by the shape of the aedeagus.

Figures

Figs 251 - 255: Aedeagi of Scaphisoma; (251, 252), S. trilobum nov. sp., in dorsal and lateral views, scale = 0.3 mm; (253, 254), ditto, internal sacs, in dorsal and lateral views, scale = 0.15 mm; (255), S. trimaculatum nov. sp., aedeagus in dorsal view, scale = 0.3 mm.

Figs 256 - 258: Scaphisoma; (256), S. trimaculatum nov. sp., aedeagus in lateral view, scale = 0.3 mm; (257), S. bicolor nov. sp., antennomeres II-XI, scale = 0.1 mm; (258), S. lienhardi nov. sp., antennomeres III-XI, scale = 0.1 mm.