Scaphisoma furcatum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1422-1423

5.2.19.8. Scaphisoma furcatum nov.sp. ( Figs 201-203 )

T y p e m a t e r i a l: Holotype 6: Mindanao Eagle Centre, 1100 m Baracatan north slope of Mt. Apo 5.VIII.1985 M. Sakai (EUMJ). Paratypes: Mindanao, with the same data as the holotype but 6.VIII.1985, 1 6 (MHNG); Mindanao, Davao Prov., 25 km NW of New Bataan, 1200 m, 20- 22.V.1996, leg. Bolm, 1 6 (SMNS); Luzon, Benguet Prov., Pellwell Ck., Camp Two, 14.VI.1977, leg. M. Satô, 1 6 (MHNG).

Description: Length 1.35-1.50 mm, width 0.95-1.0 mm. Body dark brown, pronotum slightly darker than elytra, apical abdominal segments and appendages light brown. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 15/6: IV 32/7: V 45/8: VI 45/9: VII 53/13: VIII 46/9: IX 57/12: X 52/11: XI 73/13. Pronotum on prevailing surface very finely and fairly densely punctate, punctation on basomedian area fairly coarse and dense; lateral contours evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae impunctate. Exposed tip of scutellum minute. Elytra with lateral margin carinae visible in dorsal view, apical margins truncate, inner apical angles not prominent, situated in level with outer angles; sutural margin not raised, sutural striae shallow, starting posterior basal fourth of sutural length, distinctly punctate; adsutural areas narrow, flat, densely and coarsely punctate, lateral and epipleural striae distinctly punctate. Elytral disc with punctation very dense and coarse, puncture intervals mostly smaller than puncture diameters, becoming finer and less coarse apically, punctures smaller than puncture intervals near apices; small humeral areas impunctate. Hind wings fully developed. Hypomera and mesanepisterna with strigulate microsculpture. Mesepimera about three times as long as intervals to mesocoxae, four times as long as wide. Metaventrite with strigulate microsculpture and very finely, sparsely punctate, middle slightly convex, lacking mesal or apicomesal impressions or striae; antecoxal puncture rows absent. Submesocoxal areas 0.05-0.06 mm, about as third of intervals to metacoxae, submesocoxal lines convex, almost impunctate. Metanepisternum convex, narrowed anteriad, inner margin straight, rounded near apical angles, impressed below margin of metaventrite. Tibiae straight. Abdomen with strigulate microsculpture, very finely punctate, submetacoxal areas 0.05-0.06 mm, about as half of intervals to apical margin of sternite 1, submetacoxal lines convex, distinctly punctate.

M a l e: Protarsomeres 1 to 3 distinctly widened. Aedeagus ( Figs 201-203 ) 0.63-0.75 mm long, symmetrical, strongly sclerotized. Median lobe with apical process much shorter than basal bulb, strongly inflexed, near tip perpendicular to axis of median lobe, at tip obliquely truncate (lateral view). Articular process robust, not prominent. Parameres wide in basal section, abruptly narrowed anterior mid-length, with apical halves arcuate in dorsal view, sinuate in lateral view. Internal sac complex, with robust basomedian tooth, two long sinuate admesal sclerites, scale-like, denticulate and baculiform structures.

E t y mo l o g y: The species epithet is a Latin adjective referring to the aedeagus with long, fork-like sclerites.

C o m p a r a t i v e n o t e s: This species is characterized by the peculiar form of the parameres. It may be readily distinguished from its Asian congeners, S. furcillatum described below excepted, by the elytra with strongly shortened sutural striae, the discal punctures to part larger than puncture intervals, the metaventrite lacking antecoxal puncture rows, and the microsculptured hypomera, in combination.

Figures

Figs 201 - 206: Aedeagi of Scaphisoma; (201, 202), S furcatum nov. sp., in dorsal and lateral views, scale = 0.2 mm; (203), ditto, internal sac, scale = 0.1 mm; (204, 205), S. furcigerum nov. sp., in dorsal and lateral views, scale = 0.2 mm; (206), ditto, internal sac, scale = 0.2 mm.