Scaphisoma furcillatum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1424-1425

5.2.19.10. Scaphisoma furcillatum nov.sp. ( Figs 14 , 207, 208 )

T y p e m a t e r i a l: Holotype 6: Mindanao, 1.-3.May Misamis occ. 1700 m Don Victoriano Bolm lgt., 1996 (SMNS). Paratypes: Mindanao, with the same data as the holotype, 2 66, 1 ♀ (SMNMS, MHNG); Mindanao, 25 km NW of Zamboanga, 800 m Camp Susana, 28-30.IV.1996, leg. Bolm, 1♀ (SMNS).

D e s c r i p t i o n: Length 1.18-1.30 mm, width 0.80-0.90 mm. Body dark brown to blackish, apical abdominal segments and appendages light brown to ochreous. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 14/6: IV 23/6: V 54/7: VI 45/7: VII 52/10: VIII 45/7: IX 55/10: X 50/10: XI 65/12. Pronotum on prevailing surface finely and fairly densely punctate, punctation on basomedian area fairly coarse and dense; lateral contours evenly rounded, lateral margin carinae hardly visible in dorsal view, lateral striae punctate. Exposed tip of scutellum fairly large. Elytra with lateral margin carinae distinct in dorsal view, apical margins rounded, inner apical angles not prominent, situated in level with outer angles; sutural margin not raised, sutural striae shallow, starting posterior basal fourth of sutural length, distinctly punctate; adsutural areas narrow, flat, densely and coarsely punctate, lateral and epipleural striae distinctly punctate. Basal halves of elytra with discal punctation very dense and coarse, puncture intervals mostly smaller than puncture diameters, punctation becoming finer and less coarse apically, puncture diameters smaller than puncture intervals near elytral apices. Hind wings fully developed. Hypomera and mesanepisterna not microsculptured. Mesepimera about four times as long as intervals to mesocoxae, about twice as long as wide. Metaventrite without obvious microsculpture and very finely, sparsely punctate, middle hardly convex, with very shallow apicomesal impressions; antecoxal puncture rows present, hardly impressed. Submesocoxal areas 0.03 mm, about as third of intervals to metacoxae, submesocoxal lines convex, distinctly punctate. Metanepisternum convex, narrowed anteriad, inner margin straight in middle part, rounded near angles, impressed below margin of metaventrite. Tibiae straight. Abdomen with strigulate microsculpture, very finely punctate, few distinct punctures on intercoxal process of sternite 1; submetacoxal areas 0.03 mm, about as fourth of intervals to apical margin of sternite 1, submetacoxal lines barely convex, finely punctate.

M a l e: Protarsomeres 1 to 3 distinctly widened. Aedeagus ( Figs 207, 208 ) 0.67-0.73 mm long, symmetrical, strongly sclerotized. Median lobe with dorsal branches of apical process very short, ventral branch robust, much shorter than basal bulb, obliquely inflexed, tapering toward tip (lateral view). Articular process robust, not prominent. Parameres wide in basal section, sinuate in dorsal view mid-length, each bearing large membranous lobe, apical halves converging, bent at mid-length and narrowed in apical section in lateral view. Internal sac with robust apicomedian rods joined mesally, with large basal vesicle bearing minute denticles, lacking basal sclerotized structures.

E t y m o l o g y: The species epithet is a Latin adjective referring to the presence of aedeagal fork-like sclerites.

C o m p a r a t i v e n o t e s: This species resembles S. furcatum, though it lacks hypomeral microsculpture and has punctate humeral areas of the elytra. The species is characterized by its aedeagal features, in particular by the shape of the parameres seen in dorsal view, in combination with the internal sac bearing two robust apical rods joined mesally and lacking basal sclerites.

Figures

Figs 14 - 21: Habitus of Scaphisoma; (14), S. furcillatum nov. sp.; (15), S. inopportunum nov. sp.; (16), S. lunabianum nov. sp.; (17), S. inexspectatum nov. sp.; (18), S. luteomaculatum PIC; (19), S. luteopygidiale (PIC); (20), S. maculosum nov. sp.; (21), S. mindanaosum PIC. Scale: 1.0 mm.

Figs 207 - 211: Scaphisoma; (207, 208); S. furcillatum nov. sp. aedeagus in dorsal and lateral views, scale = 0.2 mm; (209, 210), S. glabrellum nov. sp., aedeagus in dorsal and lateral views, internal sac extruded, scale = 0.2 mm; (211), ditto, apical part of male sternite 6, scale = 0, 2 mm.