Scaphisoma bilobum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1429-1430

5.2.20.1. Scaphisoma bilobum nov.sp. ( Figs 216-218 )

T y p e m a t e r i a l: Holotype 6: Philippines: Leyte Lake Danao. 1991 forest edge, 500 m leg. Schawaller et al. / 19.2. - 8.3. (SMNS). Paratypes: with the same data as the holotype, 3 66 (SMNS, MHNG); Leyte, Visca N Baybay, 100-200 m, 21.II.91, leg. W. Schawaller et al., 1 6, 1♀ (MHNG).

Description: Length 1.70-1.75 mm, width 1.17-1.22 mm. Head and body reddish-brown, apical abdominal segments and appendages light brown to yellowish.

Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 11/6: IV 38/6: V 60/7: VI 46/7: VII 55/12: VIII 43/8: IX 57/10: X 52/12: XI 58/12. Pronotum very finely and fairly densely punctate, punctation on basomedian area hardly coarser than in middle; lateral contours evenly rounded, lateral margin carinae exposed in dorsal view, lateral striae punctate. Exposed tip of scutellum fairly large. Elytra with lateral margin carinae entirely distinct in dorsal view, apical margins truncate, inner apical angles not prominent, situated somewhat posterior level of outer angles; sutural margin not raised, sutural striae fairly deep, starting at base, impunctate, weakly converging from level of scutellum to apex; adsutural areas flat, densely and very finely punctate, lateral striae distinctly punctate. Discal punctation very fine and fairly sparse, puncture intervals smaller than puncture diameters. Hind wings fully developed. Hypomera and mesanepisterna not microsculptured, smooth. Mesepimera about 1.5 times as long as intervals to mesocoxae, about three times as long as wide. Metaventrite lacking mesal stria, without obvious microsculpture, very finely and sparsely punctate between mesocoxae and on lateral areas, with fairly coarse and dense punctation on impressed apicomedian area, area between mesocoxae distinctly convex; antecoxal puncture rows present, dense, weakly impressed. Submesocoxal areas 0.04 mm, about as fourth of intervals to metacoxae, submesocoxal lines convex, coarsely punctate. Metanepisternum convex, not narrowed anteriad, inner margin straight, rounded at angles, impressed below margin of metaventrite. Protibiae straight, mesotibiae and metatibiae slightly curved. Abdomen with strigulate microsculpture, very finely punctate, coarse and dense punctures on basomedian area of sternite 1 and along submetacoxal lines excepted; submetacoxal areas 0.05 mm, intervals to apical margin of sternite 1 about 2.5 times as long as submetacoxal areas, submetacoxal lines convex.

M a l e: Protarsomeres 1 to 3 distinctly widened. Aedeagus ( Figs 216-218 ) 0.86-0.97 mm long, asymmetrical. Basal bulb moderately sclerotized, expanded apically to overlap basal part of apical process of median lobe. Latter trifid, with dorsal branches very narrow. Ventral branch of apical process asymmetrical, strongly inflexed and bent at apex in lateral view, narrowed toward mid-length and in apical half almost evenly wide in dorsal view. Parameres wide, lobed ventrally, each bearing small mesal lobe, overlapping apically. Internal sac complex, with dense denticular structures and few larger teeth in apical section.

E t y m o l o g y: The species epithet is a Latin adjective meaning two-lobed.

C o m p a r a t i v e n o t e s: This species differs drastically from its congeners by the aedeagal characters, S. duplex and S. nishikawai excepted. Externally it is very similar to S. spapuloides described below, see under the latter.

Figures

Figs 212 - 218: Aedeagi of Scaphisoma; (212, 213); S. jankodadai nov. sp., in dorsal and lateral views, scale 0.2 mm; (214), S. luzonicum PIC, in dorsal view, scale = 0.2 mm; (215), ditto, internal sac, scale = 0.1 mm; (216, 217), S. bilobum nov. sp., in dorsal and lateral views, scale 0.2 mm; (218), ditto, internal sac, scale = 0.1 mm.