Scaphisoma dissymmetricum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1405-1406

5.2.15.1. Scaphisoma dissymmetricum nov.sp. ( Figs 155, 156 )

T y p e m a t e r i a l: Holotype 6: Palawan (central) Sabang, trail to Underground River, sea level, 30.XI.95 I. Löbl, fungi on log (MHNG). Paratypes: with the same data as the holotype, 1 6 (MHNG); Palawan, Cabayugan near Lion’s Cave, sea level, 1.XII.1995, leg. I. Löbl, 1♀ (MHNG). Description: Length 1.05-1.10 mm, width 0.73-0.75 mm. Head and body reddish-brown, apical abdominal segments and appendages lighter. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 12/6: IV 20/5: V 30/6: VI 30/6: VII 38/14: VIII 29/10: IX 38/13: X 38/12: XI 50/14. Pronotum fairly coarsely, densely punctate, with lateral margins evenly rounded, lateral margin carinae barely exposed in dorsal view, lateral striae punctate. Minute tip of scutellum exposed. Elytra with lateral margin carinae hardly visible in dorsal view, apical margins rounded, inner apical angles not prominent, situated about in level with outer angles; sutural margin not raised, sutural striae deep, hardly bent at base and not extending along basal margin, parallel except in apical third, distinctly punctate; adsutural areas flat, densely and finely punctate, with punctures smaller than those on elytral disc, lateral striae finely punctate. Elytral disc with punctation dense and fairly coarse, punctures fairly well delimited, punctures intervals mostly about twice to four times as large as puncture diameters. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera twice as long as intervals to mesocoxae, about four times as long as wide. Metaventrite without mesal stria or impressions, with strigulate microsculpture except near lateral margins, punctation sparse and very fine. Anterolateral surface of metaventrite not microsculptured; antecoxal puncture rows absent. Submesocoxal areas 0.02-0.03 mm, about as fifth to fourth of intervals to metacoxae, submesocoxal lines parallel, distinctly punctate. Metanepisternum flat, hardly narrowed anteriad, inner margin almost straight, suture shallow. Tibiae straight. Abdomen with strigulate microsculpture, very finely and sparsely punctate. Sternite 1 with submetacoxal areas 0.04-0.05 mm, about as half of intervals to apical margin, submetacoxal lines convex, coarsely punctate.

M a l e: Protarsomeres 1 to 3 hardly widened. Aedeagus ( Figs 156 , 157 ) 0.41-0.45 mm long. Median lobe slightly asymmetrical, moderately sclerotized. Apical process shorter than basal bulb, inflexed, gradually narrowed, with dorsal lamina weakly sclerotized, rounded at tip. Parameres asymmetrical, with membranous ventral lobes; right parameres shorter than left, in apical section weakly bent and about as wide as near base. Left parameres strongly narrowed posterior mid-length and strongly bent (dorsal view). Internal sac with denticulate rod, several fairly long apical spine-like denticles and bulbous basal part covered by very fine scale-like and denticular structures.

Etymology: The species epithet is a Latinized adjective referring to the asymmetrical aedeagus.

Comparative notes: The species may be distinguished from the allied S. solutum by the length ratios of the antennomeres III and IV (antennomere IV is only slightly longer than III in the latter species) and the symmetrical apical process of the median lobe with blunt apex in dorsal view.

Figures

Figs 151 - 156: Aedeagi of Scaphisoma; (151), S. iridescens nov. sp., aedeagus in dorsal view, internal sac extruded; (152), ditto, paramere in dorsal view, scale = 0.2 mm; (153, 154), S. werneri nov. sp., aedeagus in dorsal and lateral views, scale = 0.2 mm; (155, 156), S. dissymmetricum nov. sp., aedeagus in dorsal and lateral views, scale = 0.1 mm.

Figs 157 - 161: Aedeagi of Scaphisoma in dorsal and lateral views; (157 - 159), S. hamatum nov. sp., internal sac extruded, scale = 0.2 mm; (159), ditto, apical part of median lobe with internal sac, scale = 0.1 mm; (160, 161), S. banguiense LÖBL, scale = 0.1 mm.