Scaphisoma bicuspidatum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1387-1388

5.2.12.1. Scaphisoma bicuspidatum nov.sp. ( Figs 5 , 118, 119 )

T y p e m a t e r i a l: Holotype 6: Philippines: Luzon Mt. Makiling, 400m 19.XI.1995, I. Löbl summit road, litter (MHNG). Paratypes: with the same data as the holotype, 2 66, 5 ♀♀ (MHNG); Lagunas Prov., Mt. Makiling, 4 km Los Banos, 8.IV.1977, leg. L. Watrous, 2♀ (MHNG); Luzon, Lagunas Prov., Mt. Makiling, above Mad Spring, 400-700 m, 19-22.XI.1995, leg. J. Kodada, 2 66, 4 ♀♀ (MHNG).

D e s c r i p t i o n: Length 1.45-1.55 mm, width 1.05-1.10 mm. Head and most of body blackish-brown to black. Apical fifth to fourth of elytra yellowish, narrowly dark brown apices excepted. Apical abdominal segments, femora and tibiae reddish-brown, antennae and tarsi yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 11/7: IV 28/6: V 40/7: VI 40/6: VII 50/12: VIII 38/8: IX 48/12: X 50/ 12: XI 52/13. Pronotum finely and very densely punctate, puncture diameters to part about as puncture intervals, with lateral margins evenly rounded, lateral margin carinae partly exposed in dorsal view, lateral striae very finely punctate. Visible tip of scutellum minute. Elytra not flattened, with lateral margin carinae entirely visible in dorsal view, apical margins truncate, inner apical angles not prominent, situated posterior level of outer angles; sutural margin not raised, sutural striae fairly deep, bent at base, extended along basal margin to form basal striae almost reaching basal mid-width, parallel posterior level of scutellar tip except near apices, impunctate; adsutural areas narrow, flat, each with single puncture row, lateral striae distinctly punctate. Elytral disc with punctation fine and dense, mostly coarser than that of pronotum, with some punctures about as large as puncture intervals, punctation near bases and lateral margins similar to that on pronotum. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera about as long as intervals to mesocoxae, hardly three times as long as wide. Metaventrite not microsculptured, extremely finely punctate, few larger punctures on apicomedian area excepted. Median part of metaventrite flattened, apically somewhat impressed; antecoxal puncture rows absent. Submesocoxal areas 0.05 mm, about as third of intervals to metacoxae, submesocoxal lines convex, distinctly punctate, punctures not extended by striae. Metanepisternum flat, weakly narrowed anteriad, inner margin arcuate, suture deep. Tibiae straight. Abdominal sternite 1 lacking obvious microsculpture, very finely and sparsely punctate; submetacoxal areas 0.04 mm, about as fourth of intervals to apical margin of sternite, submetacoxal lines convex, distinctly punctate. Following ventrites with punctulate microsculpture.

M a l e: Protarsomeres 1 to 3 and mesotarsomeres 1 and 2 distinctly widened. Aedeagus ( Figs 118, 119 ) 0.71 mm long, fairly sclerotized, asymmetrical. Median lobe narrow, with strongly sclerotized dorsal valve and abruptly inflexed apical section, ostium large, dorsal, tip truncate and with acute angles in lateral view. Articular processes robust, not prominent. Parameres with strongly extended bases, widest about in middle in lateral view, with narrowly membranous inner margins seen in dorsal view. Internal sac strongly sinuate, flattened, sclerotized except in apical section, latter extruded in resting position, lacking denticular or spine-like structures.

E t y m o l o g y: The species epithet is a Latin adjective, meaning double-pointed and referring to the shape of the apex of the median lobe, seen in lateral view.

C o m p a r a t i v e n o t e s: This species possesses a unique shape of the median lobe and internal sac. It may be, however, related with S. scurrile with which it shares a large dorsal ostium, and the internal sac strongly sclerotized and in resting position extruded, lacking denticular or spinous structures.

Figures

Figs 1 - 6: Habitus of Scaphisoma; S. anomalum LÖBL (1); S. bibiranense LÖBL (2); S. bicolor nov. sp. (3); S. bicoloripenne nov. sp. (4); S. bicuspidatum nov. sp. (5); S. binaluanum PIC (6). Scale: 1.0 mm.

Figs 115 - 119: Aedeagi of Scaphisoma; (115), S elpis nov. sp., lateral view, without contours of proximal part of basal bulb, scale = 0.1 mm; (116, 117), S. alticola nov. sp., in dorsal and lateral views, scale 0.2 mm; (118, 119), S. bicuspidatum nov. sp., in dorsal and lateral views, scale 0.2 mm.