Scaphisoma scapulare, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1395-1396

5.2.13.12. Scaphisoma scapulare nov.sp. ( Figs 30, 135-137 )

T y p e m a t e r i a l: Holotype 6: Luzon: Philippines Mt. Puguis (1900m) nr. Bontoc Mountain Pv. 20.VII.1985 M. Sakai leg. (EUMJ). Paratypes: Luzon, Mountain Prov., Mt. Data Lodge, 2200-2300 m, 22-23.XII.79, L. Deharveng & J. Orousset, 1 6 (MHNG); Luzon, Mountain Prov., Mt. Data, 2250 m, 13.VII.1985, leg. M. Sakai, 1 6 (EUMJ); with the same data but 25.VII., 4 66, 6 ♀♀ (EUMJ, MHNG, BZLA); with the same data but 15.VII., 1 6 (EUMJ); with the same data but 2300 m, 26.VII., 3 66, 5 ♀♀ (EUMJ, MHNG); Luzon, Mountain Prov., Mt. Puguis, 1800- 1900 m, 17.VII.1985, leg. M. Sakai, 2 66, 6♀♀ (EUMJ, MHNG); Mt. Puguis, 2000 m, 5- 6.VI.1977, leg. M. Satô, 2 66 (MHNG); Mt. Pangao, 2400 m, 30.V.1977, leg. M. Satô, 1 6, 2♀♀ (MHNG); Luzon, Benguet Prov., Paoay, 11.VII.1985, leg. M. Sakai, 2♀♀ (EUMJ); Luzon, Bangui, 1 6 (SMTD).

D e s c r i p t i o n: Length 1.70-2.10 mm, width 1.10-1.50 mm. Head, thorax, most of elytra, and abdomen dark reddish-brown to blackish-brown. Apical fourth of elytra light, reddish. Femora reddish-brown, tibiae, tarsi and antennae lighter than femora, almost yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 15/11: IV 30/8: V 35/9: VI 47/10: VII 55/14: VIII 33/12: IX 50/16: X 50/20: XI 65/23. Pronotum finely and densely punctate, with lateral margins evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae impunctate. Exposed tip of scutellum minute. Elytra convex, with lateral margin carinae hardly visible near base or entirely concealed in dorsal view, apical margins truncate, inner apical angles not prominent, situated posterior level of outer angles; sutural margin slightly raised, sutural striae shallow, bent at base, extended along basal margins to form basal striae reaching basal mid-width or outer third of basal width, parallel along suture except near apices, very finely punctate; adsutural areas narrow, flat, each with single puncture row, lateral striae impunctate. Elytral disc with punctation very fine, similar to that on pronotum. Hind wings fully developed. Hypomera and mesanepisterna smooth.

Mesepimera about as long as intervals to mesocoxae, about four or five times as long as wide. Metaventrite not microsculptured. Median part of metaventrite flattened, without impressions, distinctly punctate, lateral parts of metaventrite with extremely fine and sparse punctation; antecoxal puncture rows absent. Submesocoxal areas 0.06-0.08 mm, about as thirds of intervals to metacoxae, submesocoxal lines convex, distinctly punctate, punctures not extended by striae. Metanepisternum flat, narrowed anteriad, inner margin rounded, suture deep. Tibiae straight. Abdomen with hardly visible or evanescent strigulate microsculpture, and very finely and sparsely punctate, except on small basomedian area of sternite 1 bearing distinct punctures; submetacoxal areas 0.05-0.07 mm, about as third to half of intervals to apical margin of sternite, submetacoxal lines convex, distinctly punctate.

M a l e: Protarsomeres 1 to 3 and mesotarsomeres 1 and 2 distinctly widened. Aedeagus ( Figs 135-137 ) 0.53-0.71 mm long, asymmetrical, strongly sclerotized. Basal bulb fairly large. Articular processes large, not prominent. Apical process of median lobe split into two laminae, with large ostium in-between. Right lamina about as long as basal bulb, inflexed, subcylindrical in dorsal view and with acute tip, left lamina short. Parameres narrowed and with membranous inner margins in apical halves, denticular at mid-length (dorsal view). Internal sac with robust, strongly sclerotized flagellum proximally bent and hook-like, dilated and shovel-like at apex; spines and scale-like structures absent.

E t y m o l o g y: The species epithet is a Latin adjective referring to the shovel-like apex of the internal sac of the aedeagus.

Co mp ar a t i v e n o t e s: The aedeagal characters suggest relationships with S. biliranense LÖBL, though the flagellum is not hook-like and the parameres are almost gradually narrowed in the latter species. This new species may be easily distinguished from S. biliranense by the larger body-size and light apical fourth of the elytra.

Figures

Figs 131 - 136: Aedeagi of Scaphisoma in dorsal and lateral views; (131, 132), S. palawanum PIC; (133, 134), S. pandanum nov. sp.; (135, 136), S. scapulare nov. sp. Scale = 0.1 mm.

Figs 137 - 141: Aedeagi of Scaphisoma; (137), S. scapulare nov. sp., apical part of median lobe with extruded internal sac, scale = 0.1 mm; (138, 139), S. tagalog nov. sp., aedeagus in dorsal and lateral views, scale = 0.1 mm; (140, 141), S. angulosum nov. sp., aedeagus in dorsal and lateral views, scale = 0.2 mm