Scaphisoma bicoloripenne, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1398-1399

5.2.14.2. Scaphisoma bicoloripenne nov.sp. ( Figs 4 , 142, 143 )

T y p e m a t e r i a l: Holotype 6: Philippines Bohol, Bilar 23.VII.1970 M. Satô leg. (MHNG). Paratypes: with the same data as the holotype, 1♀ (MHNG); Bohol, Longon, 23.VII.1970, leg. M. Satô, 2♀♀ (MHNG); Bohol, Bilar-Longon, 22.VII.1970, leg. M. Satô, 2 66, 3♀♀ (MHNG); Leyte, Visca N. Baybay, primatry forest 200-500 m, 22.II.1991, leg. W. Schawaller et al., 2♀♀ (SMNS, MHNG).

Description: Length 1.50-1.60 mm, width 1.08-1.15 mm. Head, thorax and most of elytra reddish-brown. Apical fourth of elytra light brown to yellowish. Most of abdomen reddish-brown, lighter than thorax. Abdominal apex and appendages yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 12/8: IV 28/5: V 48/7: VI 45/8: VII 52/14: VIII 34/10: IX 45/13: X 48/14: XI 58/14. Pronotum very finely and sparsely punctate, with lateral margins evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae impunctate. Exposed tip of scutellum minute. Elytra with lateral margin carinae entirely and well visible in dorsal view, apical margins weakly rounded, inner apical angles not prominent, situated about in level with outer angles; sutural margin raised, sutural striae deep, somewhat bent at base, not extending along basal margin, strongly converging toward apices, throughout distinctly punctate; adsutural areas conspicuously broad anteriad, about twice as wide at level of scutellar tip as at mid-length, flat, densely and distinctly punctate, with punctures usually somewhat smaller than those on elytral disc, appearing impunctate near apices, lateral striae finely punctate. Elytral disc with punctation dense, fairly coarse, with punctures well delimited, puncture intervals mostly about as large to twice as large as puncture diameters, punctation denser on middle part of lateral areas, becoming finer toward apices. Hind wings fully developed. Hypomera smooth. Mesanepisterna with strigulate microsculpture. Mesepimera longer than intervals to mesocoxa, about three times as long as wide. Metaventrite with distinct strigulate microsculpture, very finely and sparsely punctate except on small apicomedian area bearing dense, fairly coarse punctures and dense antecoxal puncture rows. Median part of metaventrite somewhat convex, flattened apically, apicomedian impressions or striae absent. Submesocoxal areas about 0.05 mm, almost as half of intervals to metacoxae, submesocoxal lines convex, distinctly punctate. Metanepisternum with strigulate microsculpture, hardly convex, moderately narrowed anteriad, inner suture deep, arcuate. Abdomen very finely and sparsely punctate, with distinct strigulate microsculpture; submetacoxal areas 0.07 mm, about as half of intervals to apical margin of sternite 1, submetacoxal lines convex, distinctly punctate.

M a l e: Protarsomeres 1 to 3 and mesotarsomeres 1 and 2 strongly widened. Aedeagus ( Figs 142, 143 ) 0.78-0.84 mm long, symmetrical, with basal bulb weakly sclerotized. Articular processes large, not prominent. Apical process of median lobe long and inflexed. Apical section of ventral side of apical process slightly sinuate, tip acute in lateral view, blunt in dorsal view, partly covered by weakly sclerotized dorsal valves. Parameres with complex bases, arcuate in dorsal view, widened and overlapping apically, inner margins broadly membranous. Internal sac complex, with spines and scale-like structures, mesal spines enlarged and strongly sclerotized.

E t y m o l o g y: The species epithet refers to the bicolorous elytra.

Comparative notes: This species may be easily distinguished from congeners with strongly converging sutural striae of the elytra and by the mesanepisterna and metanepisterna bearing strigulate microsculpture.

Figures

Figs 1 - 6: Habitus of Scaphisoma; S. anomalum LÖBL (1); S. bibiranense LÖBL (2); S. bicolor nov. sp. (3); S. bicoloripenne nov. sp. (4); S. bicuspidatum nov. sp. (5); S. binaluanum PIC (6). Scale: 1.0 mm.

Figs 142 - 145: Aedeagi of Scaphisoma in dorsal and lateral views; (142, 143), S. bicoloripenne nov. sp., scale = 0.2 mm; (144, 145) S. biplagiatum HELLER, scale = 0.1 mm.