Scaphisoma lunabianum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1371-1372

5.2.7.10. Scaphisoma lunabianum nov.sp. ( Fig. 17 , 89, 90 )

T y p e m a t e r i a l: Holotype 6: Lake Linau, N. slope Mt. Apo, Davao Prov., Mindanao.El. 6000ft. mossy forest. XI-2-1946 / under bark / CNHM-Philippine Zool. Exped. (1946-47) H.Hoogstrall leg. (FMNH). Paratypes: Mindanao, Davao Prov., Metran E. slope Mt. Apo, 6000 ft. 5.XI.1946, on decaying fleshy, gilled fungi, original forest, leg. F.G. Werner, 3 66, 2♀♀ (FMNH, MHNG, BZLA); Davao Prov., E. slope Mt. Kinley, 7000 ft., mossy forest, 23.IX.1946, leg. H. Hoogstrall, 1 6 (MHNG); Mindanao, Eagle Centre, Baracatan, 1100 m, north slope of Mt. Apo, 5.VIII.1985, leg. M. Sakai, 2 66 (EUMJ, MHNG); Mindanao, Mainit Hot Spring, 28.VII.1970, leg. M. Satô, 1 6, 1♀ (MHNG); Mindanao, Baracatan, 1500 m, 27-29.VII.1977, leg. M. Satô, 1 6 (MHNG).

D e s c r i p t i o n: Length 1.42-1.85 mm, width 0.92-1.18 mm. Head and most of body very dark reddish-brown to blackish-brown, elytra often slightly lighter than pronotum, each elytron with yellowish or ochreous transverse subapical band usually well delimited and about as wide as or wider than dark apical area. Subapical bands slightly oblique, with irregular anterior and posterior margins. Apical abdominal segments, femora and tibiae reddish-brown, somewhat lighter than most of body, tarsi and antennae lighter and tibiae. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 12/6: IV 18/6: V 30/6: VI 43/8: VII 47/14: VIII 40/9: IX 41/12: X 43/12: XI 53/14. Pronotum very finely and densely punctate, punctures hardly visible at 30 times magnification, lateral contours evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae very finely punctate. Exposed tip of scutellum minute. Elytra with lateral margin carinae usually not exposed in dorsal view, apical margins slightly rounded to truncate, inner apical angles not prominent, situated in level with outer angles or posterior outer angles; sutural margin slightly raised, sutural striae fairly deep, hardly bent at base, not extending along basal margins, almost parallel from level of tip of scutellum toward apical third, converging near apices, finely punctate; adsutural areas narrow, finely punctate. Lateral striae punctate. Basal fourth to third of elytral disc with punctation very fine and very dense, similar to pronotal punctation. Remaining discal punctation coarser than that on pronotum, punctures not well delimited, puncture intervals mostly about as large as to three times larger than puncture diameters. Hind wings fully developed. Ventral side of thorax lacking microsculpture, very finely punctate. Mesepimera about as long as intervals to mesocoxae and about four times as long as wide. Median part of metaventrite somewhat convex, with two distinct apicomedian impressions; antecoxal puncture rows absent. Submesocoxal areas 0.04 mm, about as fourth of intervals to metacoxae, submesocoxal lines convex, distinctly punctate. Metanepisternum flat, wide, not or barely narrowed anteriad, inner margin straight, suture deep. Tibiae straight. Abdomen with distinct strigulate microsculpture and very finely punctate. Sternite 1 with submetacoxal areas about 0.05 mm, about as fourth to third of intervals to apical margin, submetacoxal lines convex, with distinct marginal punctures.

M a l e: Protarsomeres 1 to 3 moderately widened. Aedeagus ( Figs 89, 90 ) 0.55-0.69 mm long, symmetrical, fairly strongly sclerotized. Apical process of median lobe almost as long as basal bulb, inflexed, with ventral side straight except near tip, in lateral view gradually narrowed and with acute tip, in dorsal view with blunt tip; articular processes large, prominent. Parameres narrowed from bases to mid-length, posterior mid-length widened, abruptly notched subapically in dorsal view, straight with narrowed, curved apices in lateral view, reaching beyond tip of median lobe. Internal sac tubular, without rods or flagellum, with short, weakly sclerotized denticle and scale-like structures.

E t y m o l o g y: The species epithet is an anagram of the name of binaluanum.

C o m p a r a t i v e n o t e s: This species shares most diagnostic characters with S. binaluanum PIC. It may be reliably distinguished only by its aedeagal features. The body-size of this species is unusually variable.

Figures

Figs 14 - 21: Habitus of Scaphisoma; (14), S. furcillatum nov. sp.; (15), S. inopportunum nov. sp.; (16), S. lunabianum nov. sp.; (17), S. inexspectatum nov. sp.; (18), S. luteomaculatum PIC; (19), S. luteopygidiale (PIC); (20), S. maculosum nov. sp.; (21), S. mindanaosum PIC. Scale: 1.0 mm.

Figs 89 - 94: Aedeagi of Scaphisoma in dorsal and lateral views; (89, 90), S. lunabianum nov. sp.; (91, 92), S. maculosum nov. sp.; (93, 94), S. maramag nov. sp. Scale = 0.1 mm.