Scaphisoma mirum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1375-1376

5.2.7.14. Scaphisoma mirum nov.sp. ( Figs 22, 97, 98 )

T y p e m a t e r i a l: Holotype 6: Meran, E. slope of Mt. McKinley, Davao Province Mindanao, 6000 ft; IX: 5: 46 / CNHM-Philippine Zool.Exped. (1946-47) F.G. Werner leg. / on decaying fleshy, gilled bracket fungi / original forest (FMNH). Paratypes: with the same data as the holotype, 11 66, 8♀♀ (FMNH, MHNG, BZLA); with the same data but lot 100 on polypore 39 Fomes applanatus, 1 6 (MHNG); Mindanao, Davao Prov., E. slope of Mt. McKinley, 6400 ft., 2.IX.1946, mossy forest, leg. F.G. Werner, 1♀ (FMNH); same but 22.VIII. lot 10 polypore; Mindanao, Davao Prov., E. slope of Mt. McKinley, 3200 ft., 7-8.IX.1946, beating, lot. 6 54, leg. F.G. Werner, 1 6 (FMNH); Mindanao, Davao Prov., Baclayan, E. slope of Mt. Apo, 6500 ft., 10- 13.XI.1946, original forest sifting humus, 1 6 (MHNG); Mindanao, Davao Prov., E. slope of Mt. McKinley, 3200 ft., 7-8.IX.1946, beating, lot 54, leg. F.G. Werner, 1♀ (FMNH); Mindanao, Davao Prov., Lake Linau, N. slope of Mt. Apo, 7900 ft., 2.XI.1946, mossy forest, leg. H. Hoogstraal, 1 6 (FMNH); Todaya, 30.VII.1970, leg. M. Satô, 1 6 (MHNG).

D e s c r i p t i o n: Length 1.90-2.10, width 1.28-1.40. Head, pronotum, ventral side of thorax and basal sternites dark brown to blackish. Elytra ochreous, with narrowly darkened bases, lateral margins, adsutural areas; each elytron with darkened band along anterior half of sutural stria, central, well delimited dark spot narrowed laterally and often reaching lateral margin, and darkened apical eighth, latter usually reaching apical margin. Legs and apical abdominal segments ochreous, antennae yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 15/9: IV 30/7: V 40/8: VI 60/8: VII 65/16: VIII 60/13: IX 62/16: X 58/17: XI 67/18. Pronotum very finely and conspicuously densely punctate, punctures well visible at 50 times magnification, lateral contours evenly rounded, lateral margin carinae not visible in dorsal view. Exposed tip of scutellum minute. Elytra with anterior section of lateral margin carinae exposed in dorsal view, apical margins slightly rounded, inner apical angles not prominent, situated about in level with outer angles; sutural margin not raised, sutural striae deep, bent at base, not extending along basal margins to form basal striae, parallel from level of tip of scutellum toward apices, very finely punctate; adsutural areas narrow, very finely punctate. Lateral striae impunctate. Elytral disc with punctation very fine and very dense, similar to that on pronotum, punctures comparatively well delimited, smaller than intervals between them. Hind wings fully developed. Hypomera and mesanepisterna lacking obvious microsculpture. Mesepimera about as long as intervals to mesocoxae and about four times as long as wide. Metaventrite and abdomen with strigulate microsculpture, very finely punctate. Median part of metaventrite convex, impressed apically; antecoxal puncture rows absent. Submesocoxal areas 0.03-0.04 mm, about as sixth to eighth of intervals to metacoxae, submesocoxal lines hardly convex, finely punctate. Metanepisternum flat, narrow, slightly or not narrowed anteriad, inner margin straight, suture shallow. Protibiae straight, mesotibiae and metatibiae slightly curved. Sternite 1 with submetacoxal areas about 0.04-0.05 mm, about as fifth of interval to apical margin, submetacoxal lines convex, with fine marginal punctures.

M a l e: Protarsomeres 1 to 3 and mesotarsomeres 1 and 2 strongly widened. Aedeagus ( Figs 97, 98 ) 0.94-0.96 mm long, symmetrical, strongly sclerotized. Apical process of median lobe longer than basal bulb, almost evenly narrow in dorsal view, except near apex, inflexed and gradually narrowed in lateral view, with apical section rather abruptly bent. Apex of median lobe acute, articular processes robust and prominent. Parameres slightly extending behind level of apex of median lobe, narrowed toward mid-length, widened in apical third and weakly curved posterior mid-length in dorsal view, almost straight and evenly broad in lateral view. Internal sac simple, without sclerotized parts, with covered dense denticular structures.

E t y m o l o g y: The species epithet is a Latin adjective meaning conspicuous.

Comparative notes: This species may be reality distinguished from its Philippine congeners by the elytral coloration. In addition, it differs notably by the very dense pronotal and elytral punctation and the aedeagal characters. The aedeagus is similar to that in S. stictum LÖBL, 1977, but differs drastically by the median lobe lacking a sclerotized dorsal valve.

Figures

Figs 95 - 100: Aedeagi of Scaphisoma in dorsal and lateral views; (95, 96), S. minutipenis nov. sp.; (97, 98), S. mirum nov. sp.; (99, 100), S. montivagum nov. sp. Scale = 0.1 mm.