Camisolus, Seago, Ainsley E., Leschen, Richard A. B. & Newton, Alfred F., 2015

Seago, Ainsley E., Leschen, Richard A. B. & Newton, Alfred F., 2015, Two new high altitude genera of Camiarini (Coleoptera: Leiodidae: Camiarinae) from Australia and New Zealand, Zootaxa 3957 (3), pp. 300-312 : 301-304

Camisolus gen. nov.

( Figs. 1–6 )

Type species: Camisolus ptinoides sp. nov.

Diagnosis. Body shape ( Fig. 1 ) long-legged, biconvex, similar to that of the New Zealand camiarine genera Camiarus and Camiarites ; dorsum clothed with long, semierect setae. Head without ocelli; epistomal suture present, without stem. Antennae elongate, with slender, interrupted five-segmented club bearing nearly enclosed periarticular gutters on segments 7, 9 and 10. Pronotum anteriorly rounded, with sides sinuately narrowed toward hind margin; hind angles and midline of hind margin produced; disc without impressions except six across base. Elytra complete (no abdominal terga exposed), without transverse strigae, and with deep strial impressions marked by large complete punctures. Scutellary shield transverse, reduced and completely fused to elytral bases; scutellary shield partially concealed by pronotum in dorsal view. Legs elongate and robust; tarsi 5-5- 5 in both sexes.

Description. Head. Head slightly elongate, weakly constricted behind eyes; eyes round and protuberant, coarsely facetted, 8 facets across at greatest length. Head without conspicuous microsculpture. Epistomal suture present, without median stem. Antennae ( Fig. 1 ) elongate and slender; antennomeres 7 and 9–11 weakly expanded, antennomere 8 cylindrical, scarcely reduced, ~0.9x as long as preceeding segment. Antennal insertions immediately posterior to epistomal suture, partially concealed in dorsal view by shallow ridge connecting anterior margin of eye to posterolateral corner of clypeus. Gular sutures present, well separated. Tentorium with well developed anterior and posterior arms and broad tentorial bridge, dorsal arms present, extending approximately halfway to vertex of head and not associated with tentorial pits.

Prothorax. Pronotum rounded in dorsal view, subquadrate, clothed with sparse vestiture of stiff, recumbent setae, lightly punctate at setal insertions but otherwise without microsculpture. Pronotum laterally explanate only in posterior half, widest in anterior third, with hind angles flattened and acutely produced posteriad. Hind margin of pronotum with six large, conspicuous, complete punctures. Pronotum with lateral carinae complete, exposing anterior part of hypomeron to dorsal view, with loose fringe of minute setae dorsally and ventrally; hypomeron glabrous. Notosternal sutures distinct, connecting ventrolateral edge of broad cervical opening with lateral articulations of trochantins. Prosternum short, broad, with transverse carina delimiting procoxal cavities anteriorly and with prosternal process produced posteriorly and expanded to form part of procoxal closure. Coxal cavities internally separated by prosternal process, not externally separated; incompletely closed behind by conjunction of prosternal process and blunt, tapering postcoxal process arising from hypomeron. Procoxae globular, coxal cavities triangular, weakly transverse. Trochantins exposed.

Mesothorax. Scutellary shield reduced, transverse, triangular, fused to elytral bases; tergites of pterothorax reduced to slender, transverse sclerite with median longitudinal process. Mesocoxae globular, narrowly separated by processes of meso- and metaventrites; mesocoxal cavities open, bounded laterally by mesanepisternal bases, delimited anteriorly and posteriorly by prominent carinae of the meso- and metaventrites. Mesoventrite subquadrate, weakly transverse, laterally fused to mesanepisterna. Mesepimera complete, elongate, extending posteriorly to mesocoxal cavity and anteriorly to form lateral regions of prepectus. Mesotrochantins bluntly triangular, not concealed.

Metathorax. Metaventrite transverse, not carinate, with short remnant of metathoracic discrimen bisecting metathoracic process. Metacoxae globular, subconical, extending to lateral margins of metaventrite. Metacoxal cavities shallow, narrowly separated, with separation formed anteriorly by metathoracic process, posteriorly by intercoxal process of abdominal sternum 3. Metepisternal-elytral clamp (metepisternal process; Fig. 7 ) present.

Elytra. Elytra completely fused, highly convex and tapering to a blunt point posteriorly, extending to abdominal apex, epipleura convex and enclosing lateral margins of abdomen and sternum; each elytron with nine deeply impressed striae (2nd, 3rd, 8th and 9th striae from suture incomplete in apical third) bearing large, complete, windowlike punctures; epipleuron not reaching elytral apex, delimited by low carina, with scattered deep punctures, visible in lateral view. Wings absent.

Legs. Legs long, slender, similar to those of other Camiarini. Femorae loosely covered with sparse coat of minute hairs; femoral apices with paired flanges enclosing the tibial base. Tibiae clothed with long, erect setae and armed apically with short, blunt spines. Claws simple, lightly curved, each with low basal tooth. Empodium bisetose. Tarsi 5-5- 5 in both sexes.

Abdomen. Greatest width of abdominal ventrites less than half as wide as the greatest width of elytra, bordered laterally by infolded epipleura. All abdominal tergites unsclerotized, transparent. Ventrite 1 longer than remaining ventrites, with raised median simple intercoxal process, bounded by paired, shallow metacoxal rests. Ventrites 1–5 heavily sclerotized, partially fused, with thick reinforced transverse strip at base of each ventrite. Spiracle 8 apparently atrophied, non-functional.

Figures

FIGURE 1. Camisolus ptinoides, male: dorsal habitus (length of beetle = 3.7 mm).

FIGURES 2 – 7. Camisolus ptinoides, male: (2) left mandible; (3) maxilla; (4) apex of mesofemur, dosal view, with blunt spine; (5) aedeagus, ventral view; (6) male genital segment, ventral view; (7) ventral view of meso- and metathorax, arrow shows location of metanepisternal clamp. Not to scale.