Paradelia intersecta ( Meigen, 1826 )

Michelsen, Verner, 2007, Taxonomic review of Eurasian Paradelia Ringdahl (Diptera: Anthomyiidae) with descriptions of two new species, Zootaxa 1592, pp. 1-44 : 18-19

13. Paradelia intersecta ( Meigen, 1826)

Figs. 5 , 15 , 69–74 , 75 –77.

Anthomyia intersecta Meigen, 1826: 175 .

Chortophila intersecta (Meigen) ; Stein 1916: 190.

Pegomyia (Anthomyia) intersecta (Meigen) ; Schnabl & Dziedzicki 1911: 111, plate figs. 287–288, 798. Chortophila (Nudaria) intersecta (Meigen) ; Karl 1928: 174.

Hylemyia (Nupedia) intersecta (Meigen) ; Ringdahl 1933a: 33.

Pseudonupedia intersecta (Meigen) ; Ringdahl 1959: 293; Hennig 1972: 437, plate figs. 683, 688–689, 711. Paradelia (Paradelia) intersecta (Meigen) ; Griffiths 1987: 794, figs. 956–958, 960–964. For further references and synonymy see Hennig (1972: 437) and Griffiths (1987: 794).

Description. Small (WL 3.6–5.1mm; n=100+).

Male. Head, body and appendages brownish black to black, most parts covered in dark olive grey to olive brown dusting; specimens of the summer generation often strongly suffused with dark brown dusting dorsally on head and body. Dark striping pattern more or less distinct on anterior part of mesonotum. Wing almost clear or more or less tinted with dark brown; calyptrae paler or concolorous with wing base. Haltere yellow to yellow ochre. Relatively short and narrow abdomen with a mid-dorsal dark stripe that widens into dark bands anteriorly on tergites II–IV; terminalia including sternite V shining, extensively devoid of dusting.

Upper frons more or less narrower than diameter of anterior ocellus; parafrontals usually contiguous in middle. Frontal setae 3–5 pairs on lower half of frons. Pair of interfrontal setulae usually present, but sometimes very fine. Genal setae arranged in two irregular rows. Parafacial fully as wide as basal width of fore tibia. Prealar seta shorter and weaker than p notopleural seta. Proepisternals 3–6; proepimerals 9–20. Prosternum bare. Vein C without setulae on dorsal and ventral surfaces. Fore tibia with 1 ad and 1–2 pv submedian setae, or these setae reduced or absent. Mid tibia with 1 ad, 1 pd and 1–2 p setae. Mid femur with longish pv setae on basal half arranged in multiple rows basally. Hind femur with pv setae on basal two-thirds becoming long and arranged in multiple rows basally. Hind tibia with 1 av, (2–)3 ad and 2 pd submedian setae. Tergite VI narrow, fused together with syntergosternum. Sternites I–IV ( Fig. 75 ). Sternite V ( Figs. 69, 70 , 76, 77 ) showing some variation in length and shape of outer lamellae. Hypopygium ( Figs. 71, 72 ): epandrium acutely produced on dorsal part; cerci short, strongly convex in lateral view; surstyli in lateral view separable from all other species except for P. a rc t i c a. Gonites ( Fig. 73 ) and phallus ( Fig. 74 ) unremarkable, not significantly different from those in other species of the P. intersecta infrasection.

Female. Head, body and appendages brownish black to black, most parts covered in dense, light bluish grey dusting. Frontal vitta black on upper part, dark to light yellow ochre on lower part. Mesonotum usually with a pair of narrow brownish black stripes following dorsocentral setal rows. Abdomen with a faded dark mid-dorsal stripe reaching mid-length of tergite V. Wing with a light brown tinge, calyptrae concolorous but paler; haltere light ochre yellow to pale yellow.

Antenna and palpus normal-sized. Parafrontal fully one-third as wide as frontal vitta. Parafacial fully as wide as basal width of fore tibia. Proepisternals 2–4; proepimerals 3–12. Prealar seta shorter and weaker than p notopleural seta. Vein C bare dorsally and ventrally ( Fig. 5 ). Hind tibia with (2–)3 ad setae. Distal part of oviscapt as in Fig. 15 .

Material examined. Numerous specimens from the following European countries: ANDORRA, AUS- TRIA, CZECH REPUBLIC, DENMARK, FINLAND, GERMANY, GREECE, ITALY, NORWAY, RUSSIA (Lapponia rossica; Karelia rossica), SLOVAKIA, SPAIN, SWEDEN, UNITED KINGDOM.

Additional material in ZMUC tentatively identified with the present species: GEORGIA [ ZMUC]: Bakuriani, c. 90km SE of Kutaisi, meadow at c. 1700m, 1 female 16.vii.1967 (student exp.). TURKEY [ ZMUC]: Izmir: Bergama–Kozak rd., 3 females 10–12.v.1993 (V. Michelsen). UZBEKISTAN [ ZMUC]: Tashkent (41.15N / 69.28E), along brook, 1 female 19.v.1989 (M. Barták).

Distribution. Holarctic. By far the most common and widespread species of Paradelia in Europe ( Michelsen 2004), but apparently absent from the warmest, Mediterranean parts and in North Europe replaced by P. arctica above the tree line. A few old records exist from Tunisia and eastern coastal Libya but have not been verified. The Palaearctic distribution of P. intersecta outside Europe is poorly investigated. Female specimens apparently belonging to P. intersecta have presently been seen from Georgia, Turkey and Uzbekistan, and the species is further known with certainty from China ( Fan et al. 1988). In North America ( Griffiths 1987) the species ranges from Yukon to southern California. It occurs mainly in boreal and elevated areas and is apparently absent in the eastern half of the continent.

Figures

FIGURES 1 – 5. Paradelia spp. 1. P. hedgreni, female. 2. P. hedgreni, female head. 3. P. palpata, male. 4. P. hedgreni, female, detail of left wing. 5. P. intersecta, female, detail of left wing. Same scale for 1, 3 and 4, 5 respectively.

FIGURES 11 – 15. Paradelia spp., females. Oviscapt and distal parts of oviscapt, flat mounted. 11. P. lunatifrons. 12. P. lundbeckii. 13. P. h e l l en i. 14. P. palliceps. 15. P. intersecta. Same scale.

FIGURES 69 – 74. Paradelia intersecta, male. 69, 70. Sternite V, lateral and ventral views. 71, 72. Hypopygium, caudal and lateral views. 73. Pre- and postgonites, lateral view. 74. Phallus, lateral view. Same scale for 69, 70 and 71 – 74 respectively.

FIGURES 75 – 77. Paradelia intersecta, male. 75. Sternites I – IV. 76, 77. Sternite V, ventral views, specimens from same locality in southern Sweden. Same scale for 76, 77.