Hercostomus ibericus, Naglis, Stefan & Barták, Miroslav, 2015

Naglis, Stefan & Barták, Miroslav, 2015, Dolichopodidae (Diptera) from the Iberian Peninsula, with description of three new species, Zootaxa 3964 (1), pp. 125-137 : 128

Hercostomus ibericus sp. nov.

( Figs 2 A–B)

Type material. HOLOTYPE ♂: SPAIN, Sierra Nevada, Puerto de la Ragua, 2000 m, SW, 15.VIII.2006, M. Barták ( CULSP). PARATYPES: SPAIN, 5 ♂, same data as holotype, M. Barták ( CULSP).

Diagnosis. Antenna black, first flagellomere 1.5 times as long as high; arista dorsal; face with dense greyishwhite pruinosity; lower postocular setae white; thorax and abdomen dark metallic green, shiny; legs including coxae entirely black; fore tibia without apicoventral seta; hind tarsomeres slightly broadened; lower calypter yellow, with black setae; halter yellow; wing with dark brown infuscation; R4+5 and M strongly converging before wing margin; cercus greyish-white, with brownish apical border.

Description. Male. Body length (holotype): 2.8 mm, wing length 3.0 mm. Head: frons dark metallic green, shiny, with some grey pruinosity; face with dense greyish-white pruinosity, narrowest distance between eyes about 1.5 times the distance between ocellar setae; palpus black; proboscis yellowish brown; antennal segments black; first flagellomere 1.5 times as long as high; arista dorsal, bare, inserted at 2/3 from base, apical segment 4 times as long as basal segment; lower postocular setae white. Thorax: mesonotum dark metallic green shining, with some grey pruinosity; thoracic setae black; 6 pairs of strong dc; 6–7 pairs of long ac; scutellum with 2 strong marginal setae and 2 small lateral setae; 1 strong black ppls; pleura dark metallic green-blue, with grey pruinosity. Legs: including coxae entirely black, femora with metallic green reflection, setae and hairs black. Fore leg: coxa with some strong anterior setae; femur bare; tibia with pair of strong ad/pd setae at 1/3, 1 strong pd seta at 2/3, 2–3 small pv setae, and circlet of small apical setae; relative podomere ratios: 46:52:24:9:7:6:5. Mid leg: coxa with some anterior and a strong anterolateral setae; femur with strong anterior preapical seta; tibia with row of 4 strong ad setae; 2 strong pd setae at 1/5 and 2/3, 1 strong av seta at 2/3, and circlet of strong apical setae; relative podomere ratios: 67:77:33:19:14:10:8. Hind leg: coxa with strong lateral seta; femur with strong anterior preapical seta; tibia with 3 pairs of strong ad/pd setae at about 1/5, 1/2 and 3/4, and with circlet of strong apical setae; hind tarsomeres slightly broadened and flattened; relative podomere ratios: 72:89:25:29:20:13:9. Wing: brown infuscated, dark brown in anterior half between costa and M, veins dark brown; basal section of M shorter than distal section; R3+4 and M strongly convergent in apical third; M joining costa anteriad of apex; CuAx ratio: 0.5; RMx ratio: 4.2; lower calypter whitish, with black setae; halter stem brown, knob yellow. Abdomen: dark metallic green shining, with black hairs and setae; tergite 7 dark brown, tergite 8 metallic green. Hypopygium ( Figs 2 A–B): epandrium dark brown; cercus greyish-white, apical border brownish infuscated; hypandrium pale white, epandrial lobes and surstyli brown. Cercus twice as long as wide, with claw-like subapical seta, and with setae on ventral border as long as diameter of cercus; apv lobe with long, projecting basoventral seta and 2 small apical setae. Female: unknown.

Etymology. The name refers to the Iberian Peninsula where the species was found.

Remarks. The new species belongs to Hercostomus Group IV according to Stackelberg (1933), with black femora and white or yellow postocular setae. In the Palaearctic key ( Negrobov & Nechay 2009), H. ibericus runs to H. albibarbus Negrobov, 1976 described from Mongolia. Both species can be separated by the characters given in the key below.

Figures

FIGURE 2. Hercostomus ibericus sp. nov.: A, cercus, lateral; B, hypopygium lateral (cercus removed). Aed = aedeagus; Apvl = apicoventral epandrial lobe; Hyp = hypandrium; Dsur = dorsal arm of surstylus; Vsur = ventral arm of surstylus (scale bar = 0.1 mm).