Tasmanicosa hughjackmani, Framenau, Volker W. & Baehr, Barbara C., 2016
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Tasmanicosa hughjackmani sp. nov.
Wolverine wolf spider
Type data. Holotype. Male, Epsom, Londonderry Way [37°42'37”S, 144°18'11”E, Victoria, AUSTRALIA], 25 December 2009, V.W. Framenau ( WAM T 141161). GoogleMaps
Paratypes. 2 males, data as holotype (WAM T 100132), 1 female, data as holotype (WAM T 141162).
Other material examined. 44 males, 22 females and 10 juveniles in 49 records (Appendix B).
Etymology. The specific epithet for this wolf spider species honours the Australian actor Hugh Jackman, who played Wolverine in the X-Men film series, for his extraordinary artistic skills and more so for his numerous philanthropic activities. Wolf spiders are, of course, much more remarkable than Wolverines; for example, they are best caught by the fearless at night by spotlighting their sparkling green eyes, can orientate using polarised light even in the absence of direct sunshine or moonlight (e.g. Papi & Tongiorgi 1963; Dacke et al. 2001), can fly ( Richter 1970), use multimodal (visual, chemical, percussive) communication (e.g. Hebets 2004), their mothers carry their eggs and subsequently often hundreds of young on their back (e.g. Humphreys 1976b), and they can starve without food for more than a year ( Anderson 1974).
Diagnosis. The males of T. hughjackmani differ from all other species in the genus by the strongly S-shaped ridge of the tegular apophysis that forms a deep notch in the tegular apophysis ( Fig. 3N View FIGURE 3 ). Female genitalia ( Fig. 4C View FIGURE 4 ) most closely resemble those of T. godeffroyi ( Fig. 4E View FIGURE 4 ), but the medium septum is apically much wider (i.e. wider than the posterior transverse part) than in that species, in which the anterior part of the medium septum is as wide as the posterior transverse part.
Description. Male (based on holotype, WAM T 141161).
Total length 13.6.
Prosoma. Length 8.2, width 6.3; carapace reddish-brown with genus-specific Union-Jack pattern and distinct irregular lateral and smooth median light bands ( Fig. 14A View FIGURE 14 ); sternum dark brown and covered with black setae ( Fig. 14C View FIGURE 14 ).
Eyes. Diameter of AME 0.35, ALE 0.26, PME 0 63, PLE 0.84.
Chelicerae. Dark brown, with an elongated patch of yellowish-golden setae frontally.
Labium. Brown, with light brown anterior rim ( Fig. 14C View FIGURE 14 ).
Endites. Brown, apically yellow-brown ( Fig. 14C View FIGURE 14 ).
Legs. Femora yellowish-brown, patellae, tibiae, metatarsi and tarsi brown; venter of coxae dark brown, venter of patellae and tibiae apico-ventrally dark brown; legs overall covered with silvery setae. Opisthosoma. Length 8.2, width 5.1; dorsally with folium pattern bordered with light lines and patches ( Fig. 14A View FIGURE 14 ); venter black ( Fig. 14C View FIGURE 14 ).
Pedipalps. Cymbium with dense layer of silvery setae; tip with three macrosetae ( Figs 14E–F View FIGURE 14 ); terminal apophysis with strongly S-shaped ridge and therefore deeply notched ( Figs 14I –J View FIGURE 14 ); embolus sickle-shaped and gently narrowing towards tip; terminal apophysis broad and flat with rounded and slightly notched tip ( Fig. 14K View FIGURE 14 ).
Female (based on WAM T41162).
Total length 20.4.
Prosoma. Length 10.7, width 7.7; carapace and sternum colouration as male ( Figs 14B, D View FIGURE 14 ).
Eyes. Diameter of AME 0.39, ALE 0.28, PME 0.82, PLE 0.64.
Chelicerae, labium, endites, legs and opisthosoma. Opisthosoma length 10.9, width 7.9; otherwise as male, but labium and endites dark brown ( Figs 14B, D View FIGURE 14 ).
Epigyne. Slightly wider than long; medium septum inverted T-shaped, but anteriorly wider than posterior transverse part ( Fig. 14G View FIGURE 14 ); spermathecal heads longer than wide and only little wider than spermathecal stalks; spermathecal stalks coiled, originating medially at epigyne ( Fig. 14H View FIGURE 14 ).
Life history and habitat preferences. Habitat descriptions with records of T. hughjackmani include open forest of Pink Gum ( Eucalyptus fasciculosa ), Box-Ironbark forest or sclerophyll bushland. Here, the spiders construct a shallow burrow in leaf litter ( Fig. 1C View FIGURE 1 ). The phenology of T. hughjackmani appears similar to that of T. godeffroyi and T. fulgor . Males have been found between October and February, with most records from December. Female activity is between September and February, with a single record in May.
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