Kamabrachys campbelli, Constant, 2023
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publication ID |
https://doi.org/ 10.5852/ejt.2023.895.2289 |
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publication LSID |
lsid:zoobank.org:pub:75CAAC73-8100-4D16-B970-4A533DBC7000 |
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DOI |
https://doi.org/10.5281/zenodo.8407974 |
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persistent identifier |
https://treatment.plazi.org/id/A46A928F-5BC0-4636-8BFF-3A3EA912DF41 |
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taxon LSID |
lsid:zoobank.org:act:A46A928F-5BC0-4636-8BFF-3A3EA912DF41 |
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treatment provided by |
Plazi |
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scientific name |
Kamabrachys campbelli |
| status |
gen. et sp. nov. |
Kamabrachys campbelli gen. et sp. nov.
urn:lsid:zoobank.org:act:A46A928F-5BC0-4636-8BFF-3A3EA912DF41
Figs 5A View Fig , 16–21 View Fig View Fig View Fig View Fig View Fig View Fig
Diagnosis
The species can be recognized by the following combination of characters:
(1) posterior margin of the lateroventral lobes of the pygofer (lvl) oblique and sinuate in lateral view ( Fig. 18A View Fig ) and tapering towards the posterior in ventral view ( Fig. 18C View Fig ), inner margin of lobes straight in ventral view ( Fig. 18C View Fig ), ventral margin rounded in caudal view ( Fig. 18E View Fig );
(2) posterior margin of laterodorsal process of gonostylus (ldg) moderately emarginate, leaving a rather widely open angle between laterodorsal process and ventral process of gonostylus (vp) in lateral view ( Fig. 18A View Fig );
(3) crest of furcate process of periandrium (cfp) only along distal portion and not merging basally ( Fig. 20D View Fig ), in lateral view, tapering in distal portion ( Fig. 20E View Fig );
(4) laterodorsal process of periandrium (ldp) tapering in distal portion and curved dorsad, not surpassing lateroventral process (lvp) laterally and reaching slightly over half length of the lateroventral process (lvp) ( Fig. 20D–E View Fig );
(5) lateroventral process of periandrium (lvp) dorsoventrally flattened on long basal portion up to strongly curved laterodorsad distal portion ( Fig. 20D–F View Fig );
(6) apicodorsal process of aedeagus (adp) moderately sinuate, rather wide and stocky distally and tapering apically, and shorter than apicoventral process (avp) ( Fig. 20A–C View Fig );
(7) posterior wing with weakly marked whitish marking along sutural margin ( Fig. 16A View Fig ).
Etymology
The species epithet is a patronym referring to the Australian entomologist Thomas Graham Campbell who collected the type series.
Type material
Holotype AUSTRALIA • ♂; [Queensland], Macrossan , [Burdekin]; [19°59′50″ S, 146°26′22″ E]; 10 Jun. 1958; T.G. Campbell leg.; “Macrossan, Burekin Q., 10 Jun 1958, T.G. Campbell”; ANIC. GoogleMaps
Paratypes AUSTRALIA • 1 ♂, 1 ♀; same collection data as for holotype; ANIC GoogleMaps • 2 ♂♂; same collection data as for holotype; RBINS GoogleMaps .
Note
“Burekin” on the labels of the type specimens, is a typo for “Burdekin” River.
Description
MEASUREMENTS AND RATIOS. LT: ♂ (n = 4): 9.84 mm (9.74–10.00), ♀ (n = 1): 11.08 mm; BV/LV: ♂ (n = 1): 3.36, ♀ (n = 1): 3.52; BF/LF: ♂ (n = 1): 1.29, ♀ (n = 1): 1.33; LP+LM/BT: ♂ (n = 1): 0.82, ♀ (n = 1): 0.81; LM/LP: ♂ (n = 1): 3.07, ♀ (n = 1): 2.96; LTg/BTg: ♂ (n = 1): 2.65, ♀ (n = 1): 2.66; LW/ BW: ♂ (n = 1): 1.74, ♀ (n = 1): 1.69.
HEAD ( Figs 16A–D View Fig , 17A–D View Fig ). Slightly narrower than thorax. Vertex brown irregularly, rather poorly variegated with black; about 3.4–3.5 times as broad as long, concave with all margins slightly carinate; anterior and posterior margins rounded, nearly parallel, with anterior margin slightly more strongly rounded in middle portion. Frons brown irregularly, rather densely variegated with black, more densely black than vertex; darker along lateral angles and dorsal margin; dorsal angles marked with a small, yellow-brown subtriangular marking; often with a slightly paler median marking in upper half of middle area of frons; about 1.3 times as broad as long, slightly convex, slightly rugulose with peridiscal carina slightly marked; upper margin of frons slightly rounded in perpendicular view of frons; lateral margin slightly incurved above lateral projection of frons, slightly rounded under the projection. Clypeus reddish brown with median line and regular oblique lines on each side of postclypeus darker, anteclypeus reddish brown with sides black along outer margin and with median pale reddish-brown line on carina; clypeus reaching mesocoxae, elongate, with median carina on anteclypeus prolongated with obsolete carina on postclypeus; base of clypeus rounded. Labium brown, reaching metacoxae, with apical segment as long as broad, acuminate, shorter and slightly more slender than penultimate. Genae yellow-brown. Ocelli absent. Antennae brown, not surpassing eye, not visible from above; scape about as long as broad, pedicel subcylindrical, elongate, narrowing towards apex.
THORAX ( Figs 16A, C View Fig , 17A, C View Fig ). About 1.2 times as broad as combined length of pro- and mesonotum. Pronotum brown irregularly variegated with black, similarly as vertex, with portion anterior to anterior carina yellowish and usually a transverse black marking in middle of disc; disc weakly wrinkled, carina parallel to anterior margin; 2 obsolete impressed points on disc usually included in black marking; paranotal lobes brown. Mesonotum brown irregularly variegated with black, slightly more densely black and generally darker than pronotum; disc weakly wrinkled, median and peridiscal carinae weakly marked; median carina stopped before scutellum. Tegulae bicolour with ventral half black and dorsal half pale brown.
TEGMINA ( Figs 16A–C View Fig , 17A–C View Fig ). Brown irregularly variegated with darker and paler black markings; anteapical transverse white line strongly narrowing in middle portion, often reduced to 2 white, elongate triangles; apex often with a narrow white marking extending along up to 4 cells; sometimes 2 incomplete transverse black bands, more basal one at basal ¼, second one at mid-length of tegmen, often only basal band just marked by a dark marking in clavus. Flat, elongate, about 2.65 times as long as broad; costal margin weakly rounded along basal fifth, then straight nodal line; apex rounded.
VENATION. Veins ScP+RA and RP separated close to base; first fork of MP very basal, at the level of ScP+RA–RP separation; first fork of CuA slightly before apex of clavus; clavus closed; Pcu and A1 fused at about ¾ of clavus length; Pcu+A1 reaching apical angle of clavus; numerous cells along posterior half of costal margin; numerous cross-veins on apical third delimitating small elongate cells.
POSTERIOR WINGS ( Figs 16A–B View Fig , 17A–B View Fig ). Pale yellow with apical ⅓ and moderately broad band along posterior margin blackish brown (apical portion darker), anal area largely grey, somewhat paler area along middle portion of posterior margin. Well developed, as broad as tegmina, about 1.7 times as long as broad, rounded apically; anal area well developed; sutural margin weakly trilobous; not reaching apex of tegmina at rest. All main veins visible from base, forked at or after nodal line and forming a few closed cells; few transverse veinlets delimiting elongate cells on apical ¼; veins A1 and A2 sometimes with 2 terminals.
LEGS ( Figs 16A–D View Fig , 17A–D View Fig ). Pro- and mesofemora and tibiae reddish brown more or less densely variegated with dark brown; pro- and mesotarsi paler than corresponding tibia but slightly darker apically; posterior legs reddish brown with apex of tibial and tarsal spines darker. Pro- and mesofemora and tibiae dorsoventrally flattened, elongate and slender; metatibiae with 3 lateral and 10 apical spines; first metatarsomere ventrally with darker pad of microsetae at internoapical angle and two rows of 5 spines. Metatibiotarsal formula: (3) 10/4/0.
MALE TERMINALIA ( Figs 18–20 View Fig View Fig View Fig ). Pygofer (Py) rather short, as high as long and oblique in lateral view ( Fig.18A View Fig ), about 2.0 times as wide as high in caudal view ( Fig. 18E View Fig ); basal apodeme (bap) well developed and sclerotized, sinuate in dorsal view ( Fig. 18A–C View Fig ); dorsal portion narrow ( Fig. 18A–B View Fig ); lateroventral lobe (lvl) projecting posterolaterally and rounded but tapering towards the anterior and posterior in dorsal view ( Fig. 18B View Fig ); in ventral view, tapering and narrowly rounded anteriorly and posteriorly, more broadly rounded laterally and with inner margin straight ( Fig. 18C View Fig ); in lateral view mostly obliquely truncate posteriorly, with posterior margin slightly sinuate along upper + / 5 and ventral 1 / 5 somewhat angularly projecting posterad, anteroventral angle more or less rounded ( Fig. 18A View Fig ); ventral portion separated from lateroventral lobe by a well-marked, rather narrow depression ( Fig. 18C, E View Fig ); ventral lamina (vlp) about 3.2 times as broad as long with anterior and posterior margins subparallel in ventral view, posterior margin slightly bisinuate with posterolateral angles rounded and weakly projecting posterad ( Fig. 18C View Fig ). Anal tube about as broad as long (An) dorsoventrally flattened ( Fig. 18A View Fig ), oval with base and apex more strongly rounded than side margins; side margins weakly tapering towards apex; apical margin slightly indented in middle ( Fig. 18B View Fig ); paraprocts (pa) at basal ¼, well developed. Gonostyli (G) 1.29 times as broad as long in ventral view ( Fig. 18C View Fig ); ventral process (vp) in ventral view subtriangular with lateral margin very weakly sinuate, then narrowly rounded apically ( Fig. 18C View Fig ); laterodorsal process (ldg) partly visible in ventral view ( Fig. 18C View Fig ), sinuate in caudal view with mediolateral angle (mla) well-marked ( Fig. 18E View Fig ), mediolateral angle projecting laterally ( Fig. 18A, D View Fig ); dorsal hook (dhg) curved in lateral view, with anterior margin angularly sinuate basally and posterior margin indentate basally ( Fig. 18A View Fig ), and apex of hook curved cephalad ( Fig. 18B View Fig ). Connective (cv) strongly developed, about 1.5 times as long as aedeagus s. lat., with anterior pair of lateral apodemes forming a moderately broad lateral lamina directed posterolaterad and broadened apically; posterior pair of apodemes ribbon-like, strongly developed laterally and sinuate ( Fig. 19A–D View Fig ). Aedeagus s. lat. moderately upcurved with aedeagus s. str. slightly surpassing periandrium posterad ( Fig. 19E–I View Fig ). Aedeagus s. str. ( Fig. 20A–C View Fig ) with apical processes about 7.6 times as long as broad at mid-length; apicoventral process (avp) smoothly tapering towards the posterior and narrowly rounded apically; apicodorsal process (adp) in dorsal view ( Fig. 20A View Fig ) rather strongly inflated and stocky, then abruptly tapering towards apex, ending in a thick point and much shorter than apicoventral process. Dorsal periandrium ( Fig. 20D–F View Fig ) with laterodorsal processes (ldp) more or less oval in cross-section and gently curved dorsad along basal ⅔, then abruptly curved with apical ⅓ directed dorsad, strongly tapering and apically pointed; lateroventral processes (lvp) dorsoventrally flattened on basal + / 5, then strongly curved laterad and strongly tapering into an apical point directed laterodorsad, slightly broadening from base to first ⅓, then gently tapering up to distal 1 / 5, projecting posterolaterally largely beyond laterodorsal processes; basomedian processes (bmp) apically truncate in dorsal view and with sides parallel in ventral view; median furcate process (mfp) with furca slightly shorter than shaft ( Fig. 19I View Fig ) and moderately upcurved in lateral view ( Fig. 20E View Fig ), dorsal crest (cfp) on distal ¾ of arms of furca, not reflexed in dorsal view ( Fig. 20D View Fig ), crest broad and rounded basally then gently tapering towards apex and with dorsal margin undulated in lateral view ( Fig. 20E View Fig ), inner and outer lateral walls of furca slightly granulose ( Fig. 20D View Fig ).
FEMALE TERMINALIA ( Fig. 21 View Fig ). Abdominal sternite VI (As VI) with sides upcurving in posteroventral view ( Fig. 21C View Fig ); posterior margin in ventral view ( Fig. 21A View Fig ) moderately projecting posteriorly on sides; median subquadrate emargination wide; subtriangular processes more or less digitiform, rather elongate, apically rounded and directed posterodorsad ( Fig. 21A, C View Fig ); emargination between subtriangular processes and lateral projection moderately deeply rounded ( Fig. 21A, C View Fig ); processes of furca of sternite VI (FAs VI) pointed, diverging, moderately developed and curved posterodorsad ( Fig. 21A, C View Fig – mostly hidden under gonapophysis IX on Fig.). Inner portion of gonocoxa VIII (Gx VIII) rather strongly curved dorsad ( Fig. 21C View Fig ); posterior margin of gonapophysis (Gy VIII) projecting rather strongly posterad and apically rounded ( Fig. 21A View Fig ). Lateroventral excavate membranous fold of gonocoxae VIII between sternite VI and gonapophysis VIII in posteroventral view widely open ( Fig. 21C View Fig ). Inner portion of gonapophysis VIII, subtriangular processes and adjacent part of posterior margin of sternite VI, and processes of furca of sternite VI (FAs VI) densely covered in short strong setae ( Fig. 21A, C View Fig ). Gonoplacs (Gp) unilobed, large and semicircular, mostly hiding the anal tube ( Fig. 21B View Fig ). Gonocoxal bases IX (Gb IX) projecting laterally ( Fig. 21A, C View Fig ). Gonapophysis IX (Gy IX) flattened laterally, strongly reflexed posterodorsad and strongly tapering apically ( Fig. 21B View Fig ). Anal tube (An) elongate and narrow, curved posteroventrad, v-shaped in cross section beyond anus.
SEXUAL DIMORPHISM. Female about 13% larger than male.
Distribution and biology
The species is currently recorded from a single location of the middle portion of eastern Queensland, near Charter Towers ( Fig. 5A View Fig ), in Macrossan. Bioregion: Einasleigh Uplands.
The five specimens of the type series were all collected on the same day during the month of June.
Host plant unknown.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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