Eurydera ocellata Kavanaugh and Rainio, 2016

Kavanaugh, David H. & Rainio, Johanna, 2016, Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar, Proceedings of the California Academy of Sciences 63 (7), pp. 201-268 : 241-243

Eurydera ocellata Kavanaugh and Rainio , sp. nov.

Figures 23–24

TYPE MATERIAL.— Holotype ( Figs. 23A–B ), a male, in CAS, labeled: “CASENT 8068800”/ “ MADAGASCAR: Province Fianarantsoa, Parc National Ranomafana , Bellevue at Telatakely , elev 1020 m 12-19 February 2002 ”/ “ 21°15.99’S, 47°25.21’E collector: R. Harin’Hala California Academy of Sciences malaise, secondary tropical forest MA-02-09C-16”/ “ HOLOTYPE Eurydera ocellata Kavanaugh & Rainio sp. n. 2015” [red label] . Paratypes (only 1): a female (in NMNH) labeled “ MADAGASCAR: Prov. Fianarantsoa, 7 km W Ranomafana , 900 m 1-9 February 1990 W. E. Steiner ”/ “at black light in montane rainforest near river and stream”/ “ PARATYPE Eurydera ocellata Kavanaugh & Rainio sp. n. 2015” [yellow label] .

TYPE LOCALITY.— Madagascar, Fianarantsoa Province, Ranomafana National Park .

DERIVATION OF SPECIES NAME.— The species epithet, ocellata , is the Latin adjective meaning having little eyes or marked with spots. The name refers to the small, faint dark spot on interval 3 within the subapical pale area of each elytron at the insertion point of the subapical discal setiferous pore.

RECOGNITION.— Size small for genus, SBL of male and female = 9.2 mm. Members of this species ( Fig. 23A ) key to Eurydera mormolycoides Coquerel (1851) in Jeannel’s (1949) key on the basis of the following combination of features: size small; head ( Fig. 24A ) with frontal furrows deeply impressed, smooth, divergent posteriorly and without a transverse impression uniting them, posterior part of frons smooth, narrow and triangular, without lateral grooves prolonged posteriorly beyond the posterior border of the eyes; mandibles short and broad, lateral border not broadly explanate, lateral margin only slightly convex; pronotum cordate, distinctly narrowed basally, lateral margins angulate at insertions of midlateral setae; elytra short, round and black, each with a reddish subapical spot tangential to the medial suture as well as subhumeral and subapicolateral reddish areas and reddish apical spines, elytral intervals moderately convex; median lobe of male genitalia without a large, angulate projection on the ventral margin of the shaft near the base. They differ from E. mormolycoides members in having: dorsal surface slightly shiny (dull in E. mormolycoides members); pronotum with lateral explanations pale testaceous (darker in E. mormolycoides members), anterior angles broadly rounded (more narrowly rounded in E. mormolycoides members) and lateral margins with shallow but distinct sinuation anterior to hind angles (sinuation absent from or less distinct in E. mormolycoides members); elytra with larger but less distinct (less contrasting with rest of elytra) pale subapical macula with black eye spot on interval three (macula extended further laterally and lateroanteriorly than in E. mormolycoides members) and apical spine about half as long as spine in E. mormolycoides members. Finally, the median lobe of the male genitalia ( Figs. 24B–C ) has a thicker shaft throughout and an apex shorter and thicker in lateral view ( Fig. 24B ) and broader and straighter in dorsal view ( Fig. 24C ) than in E. mormolycoides males (see Jeannel 1949, Fig. 492b).

GEOGRAPHICAL DISTRIBUTION.— At present, known only from the type locality.

HABITAT DISTRIBUTION.— The holotype specimen was collected in a malaise trap in secondary montane rainforest in the Talatakely area at an elevation of 1030 m. The paratype specimen was collected at ultraviolet light in secondary montane rainforest near the junction of the Namorona River and a small tributary stream.

Figures

FIGURE 23. Digital images of holotype male of Eurydera ocellata Kavanaugh & Rainio sp. nov. A. Habitus, dorsal aspect; B. Label. Scale line = 1.0 mm.

FIGURE 24. Digital images of holotype male of Eurydera ocellata Kavanaugh & Rainio sp. nov. A. Head, dorsal aspect; B–C. Aedeagus of male genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm.