Coccidae, Fallen, 1814
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publication ID |
https://doi.org/ 10.11646/zootaXa.4460.1.1 |
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publication LSID |
lsid:zoobank.org:pub:DB841017-698F-4D44-A633-461D350DC984 |
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DOI |
https://doi.org/10.5281/zenodo.5966412 |
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persistent identifier |
https://treatment.plazi.org/id/0974884C-B676-FFD7-FF6C-FA6607D7F832 |
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treatment provided by |
Plazi |
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scientific name |
Coccidae |
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Key to adult females of species of Coccidae View in CoL View at ENA in Laos
1. Stigmatic clefts each with a single stigmatic spine............................................................ 2
- Stigmatic clefts each with at least 3 stigmatic spines.......................................................... 3
2. Antenna with 4th and 5th segments each longer than wide; preopercular pores eXtending anteriorly onto thoraX; multilocular disc-pores absent from metathoraX............................................. Drepanococcus chiton View in CoL ( Figs 4 View FIGURE 4 , 5 View FIGURE 5 )
- Antenna with 4th and 5th segments each shorter than wide, or of similar length to width; preopercular pores restricted to abdomen; multilocular disc-pores present on metathoraX........................................... D. cajani View in CoL ( Figs 2 View FIGURE 2 , 3 View FIGURE 3 )
3. Marginal setae each fan-shaped.......................................................................... 4
- Marginal setae not fan-shaped............................................................................ 5
4. Preopercular pores present anterolateral to anal plates, in 3 groups on each side; legs entirely absent.............................................................................................. Paralecanium quadratum View in CoL ( Fig. 34 View FIGURE 34 )
- Preopercular pores present anterolateral to anal plates, in 2 groups on each side; legs represented by leg stubs............................................................................................. P. expansum View in CoL ( Figs 32 View FIGURE 32 , 33 View FIGURE 33 )
5. Marginal setae truncate, of 2 types (slender and long, or thick and same length as slender ones or shorter); anal plates each with apical setae truncate..................................................... Megapulvinaria maxima View in CoL ( Figs 35 View FIGURE 35 , 36 View FIGURE 36 )
- Marginal setae of a single type, tips pointed or frayed; anal plates each with apical setae sharply spinose................. 6
6. Body entirely covered with thick waX test; without white ovisac; anal plates situated on sclerotized caudal process at maturity; stigmatic spines conical or bullet-shaped................................................................... 7
- Body without obvious waXy test; sometimes with white ovisac; anal plates not situated on sclerotized caudal process; stigmatic spines normally spinose............................................................................... 11
7. WaX test star-shaped, with 7 rays; interantennal setae numbering about 7-10 pairs......... Ceroplastes stellifer View in CoL ( Figs 14 View FIGURE 14 , 15 View FIGURE 15 )
- WaX test more or less round to oval; interantennal setae numbering about 1–3 pairs................................. 8
8. Stigmatic clefts each with a large medial seta; legs poorly developed, each with tibia and tarsus fused...................................................................................................... C. rubens View in CoL ( Figs 12 View FIGURE 12 , 13 View FIGURE 13 )
- Stigmatic clefts each with medial seta, if present, never more than twice as large as other stigmatic setae; legs normally devel- oped, each with tibia and tarsus distinct.................................................................... 9
9. Ventral tubular ducts each with a swollen inner ductule, present in lateral submarginal areas of body; with about 8–10 slender marginal setae present between anterior and posterior stigmatic clefts on each side............. C. floridensis View in CoL ( Figs 11 View FIGURE 11 , 12 View FIGURE 12 )
- Ventral tubular ducts each with a narrow or filamentous inner ductule, present in submarginal area of posterior abdomen and head only; with about 2–4 slender marginal setae present between anterior and posterior stigmatic clefts on each side..... 10
10. WaX test forming a distinct horn at maturity; stigmatic spines arranged in about 6 rows by each cleft; antenna 6 segmented; legs without tibio-tarsal articulatory scleroses.................................................. C. ceriferus View in CoL ( Figs 6 View FIGURE 6 , 7 View FIGURE 7 )
- WaX test without a horn at maturity; stigmatic spines arranged in 2 or 3 rows by each cleft; antenna 7 segmented; legs with tibio-tarsal articulatory scleroses.................................................... C. cirripediformis View in CoL ( Figs 8 View FIGURE 8 , 9 View FIGURE 9 )
11. Anal plates together kite-shaped, each plate with anterolateral margin over 1.7 times as long as posterolateral margin............................................................................. Milviscutulus mangiferae View in CoL ( Figs 37 View FIGURE 37 , 38 View FIGURE 38 )
- Anal plates together approXimately quadrate, each plate with anterolateral margin similar in length to posterolateral margin 1 2
12. Ventral tubular ducts, each with a broad inner ductule, present in medial areas of head, thoraX and abdomen; in life, mature adult female with white ovisac.......................................................................... 13
- Ventral tubular ducts each with a broad inner ductule absent; or if present, either (i) present in medial area of thoraX only (e.g. Coccus celatus and C. viridis ), or (ii) mainly present in submarginal areas of body, as well as few ducts present around meso- and procoXa (e.g. Saissetia coffeae View in CoL ); in life, mature adult female without white ovisac.............................. 15
13. Dorsum with polygonal reticulations; with 4 or 5 stigmatic spines present in each stigmatic cleft................................................................................................ Pulvinaria polygonata ( Figs 41 View FIGURE 41 , 42 View FIGURE 42 )
- Dorsum without polygonal reticulations; with only 3 stigmatic spines present in each stigmatic cleft................... 14
14. Multilocular disc-pores mostly each with 7 loculi; with ventral tubular ducts of 3 types (type I: each with a broad inner ductule; type II: each with a narrow inner ductule; type III: each with a filamentous inner ductule)........ P. floccifera ( Figs 39 View FIGURE 39 , 40 View FIGURE 40 )
- Multilocular disc-pores mostly each with 10 loculi; with ventral tubular ducts of only 2 types (type III with filamentous inner ductules absent)....................................................................... P. psidii ( Figs 43 View FIGURE 43 , 44 View FIGURE 44 )
15. Submarginal band of ventral tubular ducts present........................................................... 16
- Submarginal band of ventral tubular ducts absent........................................................... 19
16. Dorsum with polygonal reticulations; anal plates each without a discal seta; in life, without H-shaped ridge on dorsum................................................................................ Parasaissetia nigra View in CoL ( Figs 45 View FIGURE 45 , 46 View FIGURE 46 )
- Dorsum without polygonal reticulations; anal plates each with a discal seta; in life, young specimens with H-shaped ridge on dorsum............................................................................................. 17
17. With ventral tubular ducts of 3 types (type I: each with a broad inner ductule; type II: each with a narrow inner ductule; type III: each with a filamentous inner ductule), present in submarginal and medial areas of thoraX and abdomen............................................................................................ Saissetia coffeae View in CoL ( Figs 47 View FIGURE 47 , 48 View FIGURE 48 )
- With ventral tubular ducts of only 1 type (type I: each with a narrow inner ductule), present in submarginal areas only..... 18
18. Marginal setae between anterior and posterior stigmatic clefts on each side numbering 17–23...... S. miranda View in CoL ( Figs 49 View FIGURE 49 , 50 View FIGURE 50 )
- Marginal setae between anterior and posterior stigmatic clefts on each side numbering 5–13.......... S. oleae View in CoL ( Figs 51 View FIGURE 51 , 52 View FIGURE 52 )
19. Dorsum of mature adult female formed of polygonal sclerotized plates; ventral tubular ducts absent........................................................................................... Eucalymnatus tessellatus View in CoL ( Figs 30 View FIGURE 30 , 31 View FIGURE 31 )
- Dorsum of mature adult female without polygonal plates; ventral tubular ducts present, of 1 or 2 types................. 20
20. Pregenital setae numbering 1 or 2 pairs; ventral tubular ducts absent from medial area of thoraX, present only in submarginal area of posterior abdomen...................................................... Coccus capparidis ( Figs 16 View FIGURE 16 , 17 View FIGURE 17 )
- Pregenital setae numbering 3 pairs; ventral tubular ducts present in medial area of thoraX............................ 21
21. Dorsal submarginal tubercles present..................................................................... 22
- Dorsal submarginal tubercles absent...................................................................... 25
22. Ventral tubular ducts in medial area of thoraX each with a broad or filamentous inner ductule......................... 23
- Ventral tubular ducts in medial area of thoraX each with a narrow inner ductule.................................... 24
23. Multilocular disc-pores mostly each with 10–12 loculi; ventral tubular ducts of 2 types (type I: each with a broad inner ductule; type II: each with a filamentous inner ductule); antenna 8 segmented........................... C. celatus ( Figs 18 View FIGURE 18 , 19 View FIGURE 19 )
- Multilocular disc-pores mostly each with 7 loculi; ventral tubular ducts of 1 type (type I: each with a broad inner ductule) only; antenna 7 segmented.................................................................. C. viridis ( Figs 28 View FIGURE 28 , 29 View FIGURE 29 )
24. Antennae each 8 segmented....................................................... C. gymnospori ( Figs 22 View FIGURE 22 , 23 View FIGURE 23 )
- Antennae each 7 segmented....................................................... C. hesperidum ( Figs 24 View FIGURE 24 , 25 View FIGURE 25 )
25. Dorsal setae tapering, sharply setose; marginal setae with pointed apices; ventral tubular ducts abundant, a continuous transverse band of ducts present on mesothoraX; legs without tibio-tarsal articulatory scleroses........ C. formicarii ( Figs 20 View FIGURE 20 , 21 View FIGURE 21 )
- Dorsal setae cylindrical, bluntly spinose; marginal setae with bifid or fimbriate apices; ventral tubular ducts scarce, a continu- ous transverse band of ducts absent from mesothoraX, a few present near each mesocoXa; legs with tibio-tarsal articulatory scleroses................................................................... C. latioperculatum ( Figs 26 View FIGURE 26 , 27 View FIGURE 27 )
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.