Nannaria daptria, Means & Hennen & Marek, 2021

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180 : 1

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scientific name

Nannaria daptria

sp. nov.

Nannaria daptria sp. nov. Figs 98 View Figure 98 , 99 Vernacular name: "The Picnicking Twisted-Claw Millipede" View Figure 99

Material examined.

Holotype: United States - Tennessee • ♂; Tennessee, Greene County, Paint Creek Corridor , Overlook Picnic Area , forest across road; 35.9777°N, - 82.8478°W; elev. 500 m; 16 June 2018; hand collected; D. Hennen leg.; VTEC MPE04156 View Materials . For detailed collection data see Suppl. material 7. GoogleMaps


Adult males of Nannaria daptria sp. nov. are distinct from other Nannaria and the nearby N. scutellaria , based on the following combination of characters: Gonopods. Gonopodal acropodite gently curving medially throughout, not straight with 90° medial bend as in N. scutellaria . Tip blunt, with small, triangular lateral and medial flanges (Fig. 98A View Figure 98 , red arrows), not simple as in N. minor . Acropodite swollen before apex (Fig. 98A View Figure 98 , red triangle). Acropodite lacking tooth-like medial flange of N. minor , or acicular medial flange of N. scutellaria . Prefemur with long, thin, medially curving prefemoral process, not laterally curving as in N. minor and N. scutellaria . Prefemoral spine prominent, claw-like, curving cephalically, not small as in N. minor . Telopodite basal zone > 1/3 length of acropodite, not < 1/3 as in N. scutellaria . Color. Tergites with orange paranotal spots (Fig. 99 View Figure 99 ). Light brown to black background. Dorsum of collum smooth with orange and white margin.


♂ holotype (VTEC, MPE04156): BL = 30.6, CW = 3.7, IW = 2.0, ISW = 0.8, B11W = 4.8, B11H = 3.4.


No known variation.


Known only from the type locality (Tennessee: Greene County, Suppl. material 7; Fig. 126 View Figure 126 ). Distribution area: N/A; status: MRE.


Nannaria daptria sp. nov. was collected from mesic, broadleaved forests composed of oak, maple, rhododendron, and hemlock, found under deciduous leaf litter in sandy soil.


This species was named for the relaxing lunch that directly preceded its discovery. The specific name is a noun in apposition derived from the Greek daptria, meaning ‘eater.’

Type locality.

United States, Tennessee, Greene County, Paint Creek Corridor, Overlook Picnic Area, forest across road; 35.9777°N, - 82.8478°W.