Afgekia Craib, Bull. Misc. Inform. Kew 1927(9): 376 (1927), emend. nov. J.Compton & Schrire

Compton, James A., Schrire, Brian D., Koenyves 3, Kalman, Forest, Felix, Malakasi, Panagiota, Sawai Mattapha, & Sirichamorn, Yotsawate, 2019, The Callerya Group redefined and Tribe Wisterieae (Fabaceae) emended based on morphology and data from nuclear and chloroplast DNA sequences, PhytoKeys 125, pp. 1-112 : 56-57

11. Afgekia Craib, Bull. Misc. Inform. Kew 1927(9): 376 (1927), emend. nov. J.Compton & Schrire

Diagnosis.

Both species of Afgekia have two pairs of callosities on the standard petal (vs. one pair in all other genera). Stipules are the longest in the tribe 10-25 mm long (vs. 3-12 mm in Sarcodum ; 5-10 mm in Endosamara ). Floral bracts are also the longest 30-45 mm (vs. 6-20 mm in Sarcodum ). Bracteoles are absent (vs. present in Padbruggea ). The oblong pods are 6-9 cm long, smooth with a velutinous indumentum (vs. 10-25 mm long, obovate or oblong, coarsely ridged and tomentose in Padbruggea ). Seeds are flattened ellipsoid or flattened orbicular 8-13 mm thick (vs. ovoid or oblong 15-30 mm thick in Padbruggea ).

Type species.

Afgekia sericea Craib.

Genus description.

Scrambling climbers to 10-20 m. Stems green becoming brown, terete, densely sericeous. Leaves with 9-17 leaflets, evergreen, finely sericeous above and densely silvery sericeous below, imparipinnate, rachis 8-25 cm long. Stipules 10-25 mm long, linear-lanceolate, persistent. Stipels 3-5 mm long, acicular. Leaflets 3-8 × 2-3 cm, ovate to elliptic, apex softly mucronate, margins finely ciliate, base cuneate or obtuse. Inflorescence an erect leafy raceme 30-70 cm long, peduncle silvery sericeous. Flowers 23-25 mm long, emerging from June - November. Floral bracts 15-35 mm long, lanceolate, apex attenuate, densely pubescent, deep pink (purple in A. mahidoliae ), caducous. Bracteoles absent. Pedicels 7-20 mm long, sericeous. Calyx 5-7 × 5 mm campanulate, green, ivory, pinkish or purple and sericeous externally, five lobed, teeth pubescent and long acuminate, upper 2 teeth 4-8 mm long, lower 3 teeth with central tooth longest 15-17 mm, laterals 7-9 mm long. Standard 15-28 × 20-25 mm, ovate-elliptic, cream suffused with pale or dark pink or purple, sometimes streaked at base, nectar guide pale or dark yellow or greenish, back of standard densely sericeous, apex acute. Callosities in two series, a small papillate pair near the base beneath a much larger corniculate pair either side of the midline. Wing petals 20-25 × 5-7 mm, deep pink or purple, slightly falcate, glabrous except for a ciliate fringe below the apex, more or less equal in length to the keel, obovate, apex acute conjoined over the keel, basal claws 3 mm long, with either one ( A. mahidoliae ) or two auricles ( A. sericea ). Keel petals 23-26 × 7-15 mm, white, sericeous externally, broadly navicular, claws 3-8 mm long, apex rounded. Stamens diadelphous, nine fused together, the vexillary one free but lying adnate to the others, all curved upwards at apex, glabrous (basally tufted in A. sericea ). Ovary sericeous, style 1-3 mm long, glabrous, (tufted apically in A. mahidoliae ) curved upwards at apex, stigma punctate. Pods 6-15 × 3-4 cm, inflated, elliptic or oblong, dehiscent, surface smooth to slightly wrinkled, velutinous, subseptate. Seeds 2-3, flattened-ellipsoid or orbicular, 15-25 × 10-14 × 8-13 mm, hilum strap-shaped 15-30 mm long. Plate 2 A–C, E )

Etymology.

Afgekia commemorates Arther Francis George Kerr (1877-1942), Irish physician and pioneering botanist in Thailand.

Figures

Plate 2. Afgekia, Sarcodum and Padbruggea. A Afgekia mahidoliae, Thailand, Sai Yok distr. Kanchanaburi, Y. Sirichamorn s. n .. B, C Afgekia sericea Thailand S. Mattapha 1158 D Sarcodum scandens Vietnam, Quang Binh Prov. Loc & Quang P 11554 E Afgekia mahidiliae Thailand, Sai Yok distr. Kanchanaburi Y. Sirichamorn s. n.. F Sarcodum scandens Vietnam, Quang Binh Prov. Loc & Quang P 11554 G Sarcodum scandens Laos, Sop Teuang, Bolikhamxai Prov. S. Lanorsavanh 1299 H, I Padbruggea filipes Thailand, Chiang Mai, Y. Sirichamorn & S. Mattapha YSM 2017 - 1.