Soyedina washingtoni ( Claassen, 1923 )

Grubbs, Scott A. & Baumann, Richard W., 2023, The Nemourinae (Insecta, Nemouridae) of the eastern Nearctic, Zootaxa 5306 (1), pp. 1-53 : 43-44

23. Soyedina washingtoni ( Claassen, 1923)

Vernal Forestfly

http://lsid.speciesfile.org/urn:lsid: Plecoptera .speciesfile.org:TaxonName:6068

( Figs. 151–156 )

Nemoura washingtoni Claassen 1923:287 . Holotype male ( Cornell University Insect Collect ), Mt. Washington, (Coös Co.), New Hampshire, USA

Nemoura washingtoniana Needham & Claassen, 1925:217 (misspelling)

Nemoura (Soyedina) washingtoni washingtoni Ricker, 1952:53

Soyedina washingtoni Illies, 1966:249

Nemoura (Soyedina) washingtoni Hitchcock, 1974:112

Soyedina washingtoni Grubbs & Baumann, 2019:236

Distribution. Canada: NB, NS, PE. USA: CT, MA, MD, ME, NH, NY, PA, VT, WV ( DeWalt et al. 2022)

Male. Macropterous. Body length 4.6–7.8 mm, forewing length 6.5–8.0 mm (n = 10). Gills absent. Cerci simple and unmodified. Paraprocts with two lobes, inner lobes lightly sclerotized and inconspicuous, outer lobes sclerotized, quadrate in shape in lateral view with anteriorly-directed flap or spur ( Fig. 155 ). Epiproct asymmetric, typical for genus ( Fig. 151 ); ventral sclerites partially recurved over abdomen, distinctly asymmetric, apically narrowly triangular ( Fig. 151‒152 ); dorsal sclerites distinctly asymmetric, appearing scaly at higher magnifications ( Figs. 152‒154 ); open apically, inner member heavily sclerotized, flanged apically, lip symmetric apically ( Figs. 151‒152 ).

Female. Macropterous. Body length 5.4–9.7 mm, forewing length 8.0– 9.5 mm (n = 16). Gills absent. Cerci simple and unmodified. Subgenital plate well-developed, subtriangular in shape; base broad, extending from posterior of seventh sternum entirely over eighth sternum and approximately ¼ over ninth sternum; terminating in broadly-rounded apex ( Fig. 156 ).

Larva. Undescribed.

Comments. Soyedina washingtoni is distributed mainly in Atlantic Canada and the northern Appalachian Mountain region south to West Virginia and westward across Pennsylvania to small Lake Erie drainages ( Grubbs & Baumann 2019).

Figures

FIGURES 151‒156. Soyedina washingtoni. 151, USA, Maine, Thorndike Brook, epiproct, ventral view; 152, USA, Maine, Thorndike Brook, epiproct apex, ventral view; 153, USA, Maryland, spring into Elk Lick Run, epiproct, lateral view; 154, USA, Pennsylvania, Fourmile Creek, epiproct, caudal view; 155, USA, Maryland, Mill Run, outer paraproct lobe, lateral view; 156, USA, Maryland, spring into Blue Lick Run, female, abdominal terminalia, ventral view. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe, vs = ventral sclerite.