Glyptoma laeviceps SHARP, 1887

Irmler, Ulrich, 2015, The neotropical genus Glyptoma ERICHSON, 1839 with descriptions of new species and a key to the species (Coleoptera: Staphylinidae: Osoriinae), Beiträge Zur Entomologie = Contributions to Entomology 65 (2), pp. 297-325 : 301-302

Glyptoma laeviceps SHARP, 1887

( Figs 2A, B , 20A )

Glyptoma laeviceps SHARP, 1887: 723

Type material examined: Lectotype: male; Panama: Volcan de Chiriqui, 2000 - 4000 ft elev. (82°38'W, 8°46'N), under bark saturated with sap, leg. Champion ( BMNH). According to SHARP (1887) two type specimens exist. I saw one male that is here designated as lectotype.

Further material: Costa Rica: Prov. Cartago , 2 males, Par. Nat. Tapanti , cloud forest litter (dry) 1650 m elev. (83°46'W, 9°43'N), 18.02.1998, leg. R. Anderson ( KNHM) ; 2 males, 2 females, Tuis , 2 km NE Rancho Naturalista (83°33'W, 9°49'N), 2501 ft elev., 15.– 16.06.1995, leg. B. Ratcliffe, M. Jameson ( KNHM, UIC) ; Prov. Puntarenas , 1 female, Las Alturas Stanford Biolog. Sta. , ca. 29 km NE San Vito, under bark, 1500 m elev., 25.05.1993, leg. J. S. Ashe ( KNHM) ; 1 male, 1 female, same region, 2 km NE Est. Biol. Alturas , upper montane/cloud forest trans. (82°50.01'W, 8°56.56'N), litter, 1520 m elev. 10.07.1999, leg. R. Anderson ( KNHM) ; 4 males, 2 females, Las Cruces , 11 km SW Estacion Biologico Las Cruces (83°01.5'W, 8°46.43'N), wet cloud forest litter, 1450 m elev., 9.07.1999, leg. R. Anderson ( KNHM, UIC) ; 1 male, 4 females, same region (82°57.58'W, 8°47.14'N), fungus covered logs, 1330 m elev., 29.05.2004, leg. J.S.Ashe, Z. Falin, I. Hinojosa ( KNHM) ; 5 males, 2 females, Monteverde , 1570 m elev., collected by Berlese, 16.05.1989, leg. J. S. Ashe, R. Brooks, R. Leschen ( KNHM, SDEI, UIC) ; 1 male, same region, Estacion Biologica (84°47.49'W, 10°18.53'N), montane forest litter, 1600 m elev., 14.06.2001, leg. R. Anderson ; 1 male, same region (84°49.08'W, 19°19.4'N), montane forest litter, 1540 m elev., 15.05.2001, leg. R. Anderson ( KNHM) ; 1 female, Corcovado National Park, Sirena Stn. lower Ollas Trail (83°35.22'W, 8°24.48'N), under bark, 5 m elev., 25.06.2000, leg. Z. H. Falin ( KNHM) ; 3 males, 1 female, San Vito, 35 km NE, nr. Las Alturas, Rio Bella Vista , Rd to Gravel Pit , dry leaf litter near river, 4300 ft elev., 22.03.1991, 23.03.1991, leg. L. Herman ( AMNH) ; 2 females, same region, 4 km S San Vito Biol Stat. Las Cruzes (82°57.36'W, 8°47.3'N), leaf litter, 110 m elev., collected by Berlese, 19.06.1998, leg. R. Anderson ( KNHM) ; 8 males, 12 females, same region, 5 km SW S. Vito (82°59.13'W, 8°47.13'N), leaf litter, collected by Berlese, 22.06.1998, leg. R. Anderson ( KNHM, UIC) ; 1 female, Las Cruces , 4000 ft elev., 01.08.1976, C. & C. Goodnight ( AMNH) ; 1 specimen, Tapanti (83°47'W, 9°42'N), 9.09.1939, leg. A. Bierig ( FMNH) ; Prov. Alajuela , 1 female, Peñas Blancas , under bark, 930 m elev., 20.05.1989, leg. J. S. Ashe, R. Brooks, R. Leschen ( KNHM) ; Prov. Heredia , 2 females, 1 male, Puerto Viejo, Selva Biol. Stat. nr P. V. de Sarapiqui, 18.02.1985, leg. L. Herman ( AMNH) ; Panama: 1 female, Capira, Cerro Campana (79°57'W, 8°44'N), forest litter, 950 m elev., collected by Berlese, 5.06.1995, leg. R. Anderson ( KNHM) ; 2 females, Darién, Cana Biol. Stat., Serrania de Pirre (77°41.6'W, 7°45.18'N), 975 m elev., 4.06.1996, leg. A.R. Gillogly ( KNHM) ; Prov. Chiriqui , 2 males, El Hato del Volcan, NW Volcán (82°40.26'W, 8°49.16'N), litter near stream, 1410 m elev., 17.05.2001 & 24.12.2001, leg. L. Herman ( AMNH) ; 1 male, 1 female, same region, 27.7 km W Hartmann's Finca (82°44.48'W, 8°52.41'N), under bark, 1580 m elev., 17.06.1996, leg. J. S. Ashe & R. Brooks ( KNHM) ; 1 male, 1 female, same region (82°44.36'W, 8°51.48'N), forest litter, collected by Berlese, 1450 m elev., 14.06.1995, leg. R. Anderson ( KNHM, UIC) ; 1 female, La Fortuna , La Reserva, 3 km W Fortuna Hwy on continental divide trail (82°12'W, 8°46'N), 1000 m elev., 8.08.1995, leg. A.R. Gillogly ( KNHM) ; 1 male, 1 female, same region, Hydro Trail (82°14'W, 8°42'N), fungusy log, 1150 m elev., 9.06.1995 and 23.05.– 9.06.1995 collected by flight intercept trap, leg. J. S. Ashe & R. Brooks ( KNHM) ; 1 female, Los Nubes, 6.7 km N Cerro Punta, P.N. Armistad (82°34'W, 8°22'N), forest litter, 2150 m elev., collected by Berlese, 15.06.1995, leg. R. Anderson ( KNHM) .

Diagnosis: The species can be easily identified by the shiny head among the similarly sized species. In contrast to G. cicatricosa MOTSCHULSKY 1857 and G. rossii spec. nov., the head of G. laeviceps SHARP, 1887 is polished and without micro-striae.

Description: Length: 3.5–3.9 mm. Colouration: black, legs dark yellow, antennae dark brown.

Head: 0.45 mm long, 0.59 mm wide; eyes large and prominent; as long as temples; temples convergent to neck in convex curve; sides in front of eyes parallel; parallel part as long as eyes; then deeply sinuate to narrow clypeus; width of clypeus one third as wide as width of head; with swelling at base of antennae; punctation extremely sparse; only with few scattered small punctures; on average, interstices between punctures four to six times as wide as diameter of punctures; few punctures on vertex larger than others; without microsculpture; surface polished.

Antennae nearly as long as head, pronotum and elytra combined, first antennomere thick and rectangular; second only half as long as first and distinctly smaller; third approximately quadrate; following antennomeres approximately as wide as third and longer than wide; penultimate antennomere slightly thicker and again approximately quadrate; last antennomere slightly longer than penultimate; all antennomeres with long setae; setae slightly longer than width of antennomeres; surface of antennomeres matt.

Pronotum: 0.71 mm long, 0.74 mm wide; approximately globular; slightly pear-shaped; sinuate in front of posterior angles, but angles indistinct and rounded; posterior edge not wider than mesonotum; without lateral margin; punctation irregular; punctures extremely large and cori- aceous; three punctures in anterior half on both sides of midline more deeply impressed than laterally; deep furrow along midline more deeply impressed in anterior half than in posterior half; surface of midline and of punctures with deep isodiametric microsculpture, matt; interstices between punctures with weaker microsculpture and more shiny.

Elytra: 0.98 mm long, 0.98 mm wide; sides smoothly curved; widest at middle; shoulders shortly denticulate; disc with two carinae on each side of suture; one in middle between suture and lateral margin; one from shoulders to posterior angles; on inner side of discal carina three indistinct irregular rows of large punctures; on each side of suture indistinct furrow without punctures; on outer side of discal carina and inner side of lateral carina with irregular row of smaller punctures; between these rows wider space without punctures; space between carinae inclusively surface of punctures with deep isodiametric microsculpture; in some parts microsculpture longitudinally reticulate; surface matt.

Abdomen at segments III to V with deep transverse groove formed by deep large punctures; posterior half of segments impunctate; surface shiny; segments VI to VIII without transverse row of large and deep punctures and surface totally shiny; only with pair of short yellow setae on both sides of midline.

Aedeagus forming nearly rectangular angle between basal and apical lobe; apical lobe long and approximately straight; acute short apex more strongly curved and pointing ventrally; inner edge of apical lobe indistinctly undulate; wider in middle than at base; with numerous sensillae in middle; thin paramere long; reaching apical third of apical lobe; in middle with undulate structure.

Figures

Figs 20: Fore body of Glyptoma laeviceps (A), G. cicatricosa (B), G. rossii (C), G. opacum (D), G. simile (E); scale bar 0.5 mm.

Figs 2–6: Aedeagus (A), antenna (B) and microsculpture of head (C) of Glyptoma laeviceps (2), G. opacum (3), G. cicatricosa (4), G. rossii (5), G. simile (6); scale bar: 0.1 mm.