Ampullaria wernei

Cowie, Robert H., 2015, The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda, Zootaxa 3940 (1), pp. 1-92 : 52

wernei

Ampullaria wernei Philippi, 1851: 19 , pl. 5, fig. 4 [1852a: 54, pl. 17, fig. 2; 1852b: 22]. Type material — lectotype ( Köhler & Glaubrecht (2006: 212, fig. 3L): ZMB 1335 (the shell figured as wernei by Philippi (1851: pl. 5, fig. 4); = holotype of minor Bourguignat); paralectotype: the shell figured by Philippi (1852a, pl. 17, fig. 2), probably MNHNCL 15962 (1 spm.). Type locality —“der weisse Nil” [the White Nile]. Distribution —Scattered reports from a large area including southern Somalia, Sudan, possibly Kenya, Lake Chad and westwards to the Niger River in Mali and southwestern Nigeria, and western Democratic Republic of the Congo ( Brown 1994: 56).

Remarks. Both figures of Philippi (1851: pl. 5, fig. 4, 1852a: pl. 17, fig. 2) were referenced by Philippi (1851: 19), although pl. 17 may not have been published until 1852, so both specimens are part of the type series of wernei Philippi. Bourguignat (1879: 32) referenced the shell illustrated in pl. 17, fig. 2 as the “échantillon type ”. However, as Bourguignat often used the term “ type ” to mean “typical” (E. Neubert pers. comm. 25 September 2014), this cannot be considered definitively as a lectotype designation, and similarly nor can the usage of Germain (1905b: 329). Bourguignat (1879: 32) considered the shell in the other illustration of Philippi (pl. 5, fig. 4) to be sufficiently distinct from that of pl. 17, fig. 2 to warrant its recognition as variety minor Bourguignat (see minor Bourguignat), the shell of pl. 5, fig. 4, as the only representative of minor, thereby becoming the holotype by monotypy. Köhler & Glaubrecht (2006: 212, fig. 3L) designated this specimen (pl. 5, fig. 4) as the lectotype of wernei , thereby rendering wernei Philippi and minor Bourguignat objective synonyms. If Bourguignat was correct in thinking the two illustrations to be of different species, unfortunately it is the specimen that he considered to be typical of wernei Philippi (pl. 17, fig. 2) that would require a new name. Billotte (1885: 107) treated minor Bourguignat as a variety of charmesiana Billotte. Kobelt (1912b: 62–63) discussed the confusion surrounding the three names and came to essentially the same conclusion that if considered distinct, the shell illustrated in pl. 17, fig. 2 would require a new name. Pain (1961: 19) listed minor Bourguignat (as “Kobelt”) as a synonym of wernei Philippi. Reports of wernei further south than the distribution above, e.g. Mozambique and Lake Albert ( Alderson 1925: 89–90), Uganda, Zanzibar, Namibia and Angola ( Connolly 1939: 553; Pain 1961: 19), are probably incorrect identifications ( Brown 1994: 56). Pila , teste Connolly (1939: 553), Pain (1961: 18) and Brown (1994: 56). Misspelled as “ werneri ” by Gaudion (1880: 42).

winkleyi

Ampullaria winkleyi Pilsbry, 1901: 189 , pl. 5, figs. 2, 3. Type material — lectotype ( Baker 1964: 168): ANSP 76011; paralectotypes: ANSP 365369 (13 spms.), UMMZ 93221 (3 spms.). Type locality —“Henzada, Burma ”. Remarks. Valid species, teste G.B. Sowerby III (1910: 62) and Preston (1915: 103). Synonym of theobaldi Hanley & Theobald, teste Prashad (1923: 589, 1925: 77), followed here.