Pila, Roding, 1798

Cowie, Robert H., 2015, The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda, Zootaxa 3940 (1), pp. 1-92 : 28

Genus PILA Röding, 1798

POMUS Humphrey, 1797: 58 . Unavailable name: introduced in a work rejected for nomenclatural purposes ( Melville & Smith 1987: 318).

Remarks. Nevill (1885: 8) incorrectly considered Nerita urceus Müller, 1774 as the type species.

PILA Röding, 1798: 145 . Type species: Helix ampullacea Linnaeus, 1758 , by subsequent designation of Dall (1904: 53).

AMPULLARIA Lamarck, 1799: 76 . Type species: Helix ampullacea Linnaeus, 1758 , by monotypy. Remarks. Nerita urceus Müller, 1774 , invalidly designated as the type species by Dall (1904: 53).

AMPULLARIUS Montfort, 1810 . Unjustified emendation of Ampullaria Lamarck, 1799 .

PACHYSTOMA Guilding, 1828: 536 . Type species: Ampullaria globosa Swainson, 1822 , by original designation ( Guilding 1828: 539).

PACHYLABRA Swainson, 1840: 339 . Unjustified replacement name for Pachystoma Guilding, 1828 . Type species: Ampullaria globosa Swainson, 1822 .

Remarks. Swainson (1840: 339) wrote “ Pachystoma Guild., already used in Ichthyology”. Pilsbry & Bequaert (1927: 172) considered Pachystoma Guilding preoccupied by “ Pachystoma Latreille, 1809 ” ( Diptera , not ichthyology), but the latter is correctly spelled Pachystomus (see also Dall 1904: 53; Neave 1940: 517; Evenhuis & Thompson 1990: 258; Sabrosky 1999: 229). The different spellings ( Pachystoma and Pachystomus ) mean that they are not homonyms (Code, Art. 56.2). There appear to have been no earlier introductions of Pachystoma (see Neave 1940: 517).

PACHYCHEILUS Broderip, 1840: 454 . Unjustified emendation of Pachylabra Swainson, 1840 . Type species: Ampullaria globosa Swainson, 1822 .

Remarks. Baker (1930: 27) considered the name an unjustified emendation and Pilsbry (1956: 33–34) considered it merely a suggestion of how to improve Swainson’s name.

POMUS Gray, 1847: 148 . Type species: Helix ampullacea Linnaeus, 1758 , by original designation ( Gray 1847: 148).

PA CH Y C H I L U S Philippi, 1851: 7. Unjustified replacement name for Pachylabra Swainson, 1840 ; junior homonym of Pachychilus Lea & Lea, 1851 ( Pachychilidae Fischer & Crosse ). Type species: Ampullaria globosa Swainson, 1822 .

TURBINICOLA Annandale & Prashad, 1921: 7 , 9. Type species— Ampullaria nux Reeve, 1856 , by original designation ( Annandale & Prashad 1921: 9).

Remarks. Synonym of Pila Röding , teste Berthold (1991: 202).

The validity of the names Pila Röding , Ampullaria Lamarck and Ampullarius Montfort were discussed by Cowie (1997b) and ruled on by ICZN (1999a).

+ adusta

Ampullaria adusta Reeve, 1856a : pl. 3, fig. 11. Type material — syntype: NHMUK 20020690 (figured by Brown (1980: fig. 23c; 1994: fig. 23c); labelled “ Zanzibar ”). Type locality —“probably from Borneo”. Distribution —endemic to Zanzibar ( Pain 1961: 22, 1963: 152).

Remarks. Nevill (1885: 10) gave the locality as “South America ”. Confirmed as African (and hence to be placed in Pila ) by Alderson (1925: 86; see also Cowie & Thiengo 2003: 80), although Pilsbry & Bequaert (1927: 176) expressed doubt about this, synonymizing it with the South American sordida Reeve. Both adusta and pilula Reeve were first published on the same plate. G.B. Sowerby III (1910: 56) and Alderson (1925: 86) synonymized them, explicitly treating adusta as the senior synonym. Valid species, teste Kobelt (1912b: 55), who placed it in Pachylabra . Placed in Pila as a form of ovata Olivier by Brown (1980: 45, 1994: 54), followed here.