AMPULLARIIDAE Gray, 1824

Cowie, Robert H., 2015, The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda, Zootaxa 3940 (1), pp. 1-92 : 9

Family AMPULLARIIDAE Gray, 1824

AMPULLARIIDAE Gray, 1824: 276 [as “Ampullariadae”]. Type genus Ampullaria Lamarck, 1799 [= Pila Röding, 1798 ]. PILIDAE Preston, 1915: 96 . Type genus Pila Röding, 1798 .

LANISTINAE Starobogatov in Starobogatov & Sitnikova, 1983: 22. Type genus Lanistes Montfort, 1810 . POMACEINAE Starobogatov in Starobogatov & Sitnikova, 1983: 22. Type genus Pomacea Perry, 1810 . AFROPOMINAE Berthold, 1991: 206 . Type genus Afropomus Pilsbry & Bequaert, 1927 . N. syn.

ANTLIPNEUMATA Berthold, 1991: 207. Unavailable name: not based on a genus-group name (Code, Art. 11.7, 29). HETEROSTROPHA Berthold, 1991: 207 . Unavailable name: not based on a genus-group name (Code, Art. 11.7, 29). NEOPOMATA Berthold, 1991: 208. Unavailable name: not based on a genus-group name (Code, Art. 11.7, 29). NERITOPOMATA Berthold, 1991: 207. Unavailable name: not based on a genus-group name (Code, Art. 11.7, 29). SAULEINI Berthold, 1991: 206 . Type genus Saulea Gray, 1868 . N. syn.

SPIROPENIATA Berthold, 1991: 207. Unavailable name: not based on a genus-group name (Code, Art. 11.7, 29).

ICZN (1999a), following Cowie (1997b: 83–88), confirmed the name Pilidae as a junior synonym of Ampullariidae and invalid.

Starobogatov & Sitnikova (1983: 22) recognized three subfamilies in Ampullariidae (as “ Pilidae ”): Pilinae, Pomaceinae and Lanistinae . The diagnoses were minimal (dextral with calcified operculum, dextral with noncalcified operculum, sinistral hyperstrophic with non-calcified operculum, respectively) but reflect the distinctive characteristics of their type genera. However, no other information was given and these subfamilies are not recognized as valid in this catalogue , as also by Bouchet & Rocroi (2005: 247).

Berthold (1991: 206) recognized two subfamilies: Afropominae (containing the genus Afropomus only) and Ampullariinae (containing all other ampullariids). Within Ampullariinae , he recognized two tribes: Sauleini (containing the genus Saulea only) and Ampullariini (containing all other species in the subfamily Ampullariinae ). This arrangement, as the most recent formal classification, was adopted by Bouchet & Rocroi (2005: 247). Within the Ampullariini Berthold (1991: 206–212) further recognized a number of named subdivisions, listed above, none of which is nomenclaturally available. Both reanalysis of the morphological data of Berthold (1991) by Bieler (1993) and molecular analysis (Hayes et al. 2009) provide little support for the systematic arrangement of Berthold. His new subfamilies and tribes are therefore listed here as synonyms of Ampullariidae and his other new family-group names listed under Ampullariidae are unavailable.