Cabeza, HANSSON & LASALLE, 2003

HANSSON, C. & LASALLE, J., 2003, Revision of the Neotropical species of the tribe Euderomphalini (Hymenoptera: Eulophidae), Journal of Natural History 37 (6), pp. 697-778 : 704-706

Genus Cabeza gen. n.

Type species: Cabeza petiolata sp. n.

Diagnosis

Flagellum in female with two funicular segments (F1 much smaller than F2) and three claval segments (figures 66, 67, 69, 70, 72), in male with one funicular and three claval segments (figures 68, 71); flagellomeres in male with very long setae (figures 68, 71); head transverse (figures 81, 83–86), narrow at vertexal level (figure 82); scrobes present and ± U-shaped (figures 74, 81, 83–86); frontal suture present (missing or very weak close to eyes), or absent (figures 81, 85), if present then smoothly curved and situated at some distance from anterior ocellus (figures 74, 83, 84, 86); malar sulcus absent; mesosoma strongly flattened dorsally; mid lobe of mesoscutum without setae; axilla separated from mesoscutum by a sulcus; scutellum with one pair of setae, not overhanging dorsellum and dorsellum hence visible (hidden under scutellum in planiscapus sp. n.); fore wing with long admarginal setae (figures 2, 73, 76–80).

Monophyly and relationships

Cabeza belongs in the Euderomphale genus group, indicated through: vertexal suture present; axilla situated entirely anterior to scuto-scutellar sulcus; scutellum transverse, with one pair of setae; presence of a transverse occipital groove and a median longitudinal occipital groove (these grooves are very strong in Cabeza , but weak in Euderomphale and Neopomphale ).

The monophyly of Cabeza is supported by following apomorphies: flagellomeres in male with very long setae; head transverse (frontal view) and narrow at vertexal level; scrobes U-shaped; mid lobe of mesoscutum without setae; fore wing with long admarginal setae on ventral surface of membrane. Additional characters by which Cabeza is separated from the other genera of the Euderomphale genus group are: malar sulcus absent (reversal); scutellum not overhanging dorsellum and dorsellum hence visible (reversal)—however, the fact that dorsellum is hidden under the scutellum in C. planiscapus , makes this character state ambiguous in Cabeza .

Description

Flagellum in female with one small and discoid anellus, two funicular segments (one small and one large) and a three-segmented clava; in male with two small and discoid anelli, one large funicular segment and a three-segmented clava. Scape narrow or enlarged; in male with ventral sense area not visible. Mandibles with two teeth. Clypeus dorsally delimited by a smoothly curved groove, groove sometimes indistinct. Scrobes ± U-shaped (U inverted); frontal suture present (missing or very weak close to eyes) or absent, if present then smoothly curved and situated at some distance from anterior ocellus. With a groove (vertexal suture) from eye to eye, passing through ocellar triangle. Occiput with a transverse groove reaching from eye to eye, also with a longitudinal median groove between occipital margin and occipital foramen. Eyes hairy. Head narrow at vertexal level.

Pronotum reduced and strongly narrowed medially, not visible in dorsal view. Thoracic dorsum flat, with weak engraved to strong and raised reticulation, some parts smooth and shiny; notauli absent. Mid lobe of mesoscutum without setae; scutellum with one pair of setae situated in posterior or anterior half of scutellum; mesoscutum and scutellum separated by a distinct groove. Axillae situated anterior to scuto-scutellar sulcus. Transepimeral sulcus distinctly curved to almost straight. Propodeum smooth and shiny, spiracular sulcus absent; propodeal callus with two setae. Fore wing with one, two or three setae on submarginal vein; costal cell as wide as base of marginal vein; with surface just below base of marginal vein strongly to weakly infuscate; speculum open below; with 1–7 admarginal setae; postmarginal vein 0.5–1.3× as long as stigmal vein (difficult to measure in baeostigma ).

Petiole 0.5–1.6× as long as wide, dorsal surface smooth or with strong irregular sculpture. Female gaster circular.

Distribution Brazil, Costa Rica.

Hosts Cabeza petiolata is associated with Ficus brenesii (Moraceae) in some way. Etymology

Named after the peculiar looking, wide heads that species of this genus have (cabeza=head in Spanish). Gender is regarded as feminine.