Libythea laius laius Trimen 1879

Kawahara, Akito Y., 2013, <strong> Systematic revision and review of the extant and fossil snout butterflies (Lepidoptera: Nymphalidae: Libytheinae) </ strong>, Zootaxa 3631 (1), pp. 1-74 : 29-30

Libythea laius laius Trimen 1879

( Figs. 6 , 18, 30, 43, 58, 109–112)

Libythea laius Trimen 1879: 337 .

Dichora laius Fruhstorfer 1898: 170 .

Libythea (Dichora) laius: Pagenstecher 1901: 12 , 13.

Libythea (Dichora) lajus [sic] Pagenstecher 1902: 3.

Libythea werneri Fruhstorfer 1903b: 112 . Shields, 1984a: 265 [= Libythea laius laius ].

Dichora werneri Hering 1921: 264 .

Dichora laius laius: Hering 1921: 267 , 268.

Libythea labdaca laius: Pinhey 1949: 76 , pl. 11, Fig. 2 .

Libythea (Dichora) labdaca f. laius: Pinhey 1949: 94 .

Libythea (Dichora) laius laius: Peters 1952: 90 .

Libythea labdaca f. laius: Swanepoel 1953: 276 , pl. 10, Fig. 27 .

Libythea laias [sic] Ehrlich 1958: 314.

Libythea lajus [sic] Puységur 1967: 25.

Libythea labdaca laius: Cooper 1973: 106 , pl. 32, Figs. 111, 111a .

Libythea laius laius: Sevastopulo 1974: 131 .

Libythea labdaca: Williams 1994: 154 , pl. 10, Figs. 102a, b [misidentification].

Libythea (Jera) labdaca laius: Pennington 1994: 125 , pl. 93.

Diagnosis. Libythea laius laius is similar to L. l. lepitoides and L. l. tsiandava but it differs from the latter subspecies because the apical orange mark in forewing discal cell is clearly separated from the orange mark in cells M 3 and CuA 1.

Type Specimens Examined. Libythea laius Trimen 1879 , PARATYPE: ♀, Red label without writing. Printed white label, ‘NATAL/D’Urban dist.’ Printed white label, ‘31.’ Hand written white label: ‘ L. laius / Trimen CoType from Trimen.’ Printed red label, ‘ PARATYPE.’ Printed white label with black lined border: ‘ Libythea laius ♀ / Trimen 1879 / det. A. Y. Kawahara 2002’ ( BMNH).

Other Specimens Examined. (70♂, 42♀): KENYA: Amani, Usambara : 2♂, II-1950, Van Someren ( AMNH) . Chepalungu : 1♂, VII-1950, Van Someren ( AMNH) . Meru: 1♂, 8-I-1943 ( AYK) . Mombass 1♂, 3♀, 25-27-II-1912, S. A. Neave ( BMNH) . Nairobi : 1♂, 26-IV-1942, N. Mitton ( USNM) ; 1♀, III-1956, R. Carcasson ( USNM) . Near Wangi, Coast of Mainland , 3♂, 3♀, 21-II-1912, S. A. Neave ( BMNH) . Ngong : 1♂, IV-1936, Van Someren ( AMNH) . Saquana River : 1♂, 14-VI-1950, Gregson ( USNM) . MALAWI: Cholo , 2700 ft [823 m]: 2♂, 1♀, 8-10-VIII-1927, R. C. Wood ( CMNH) ; 2♀, 10-X-1927 ( AMNH) ; 11♂, 15♀, 1927- 1929 ( CMNH) . Mlanje , 1♂, 3- I, S. A. Neave ( BMNH) . Mlanje Boma , 400 ft [122 m], 3♂, 1♀, 26-30-IV, 3-5- V-1910, S. A. Neave ( BMNH) . Side of Mt. Mlange, Side of Mt. Mlange , 3000-4000 ft [914-1220 m], 1♀, 3- XII-1913, S. A. Neave ( BMNH) . Zomba, 1♂ ( BMNH) ; 6000ft [1829 m]: 5♂, 4♀, H. Barlow ( USNM) . MOZANBIQUE: 10 mi [16.09 km] west of Vila Machado : 1♂, 14-III-1958, E. S. Ross and R. E. Leech ( CAS) . Xiluvo : 1♂, 1♀, 23-IV-1971, D. M. Kroon ( MGCL) . SOUTH AFRICA: Baskee : 1♂, 7-VIII-1947, G. C. Clark ( CUIC) . Cape Province: 1♂, 28-III-1960 W. Teare ( MGCL) . Dukuduku : 1♂, 17-III-1952, A. S. Aiken ( AMNH) . Durban : 1♂, 3-III-1962, G. C. Clark ( LACM) . Emboyti : 2♀, 20-I-1952, A. S. Aiken ( AMNH) ; Malta : 1♂, III-1957, W. Teare ( MGCL) ; 3♂, 28-XII-1970, J. C. McMaster ( MGCL) . Malta Forest : 1♀, 31-XII-1967, D. A. Swanpoel ( MGCL) . KwaZulu-Natal, Umkomaas : 1♂, 20-IV-1911 ( BMNH) ; 1♂, 1♀, 8-9-II-1913, L. G. Higgins ( BMNH) . Mount Ubatuba , 1♂, 1♀, 27-II-1951, A. S. Aiken ( AMNH) . Port St. Johns : 1♂, 29-IV-1965 J. C. McMaster ( MGCL) ; 1♂, 10-I-1977, D. M. Kroon ( MGCL) . Transkei : 1♀, 11-IX- 1966, C. D. Quickelber ( MGCL) . Transvaal : 1♂, XII-1958, W. Teare ( MGCL) ; 1♂, 10-III-1962, W. Teare ( MGCL) ; 1♀, 28-IV-1962, W. Teare ( MGCL) , 4♂, 12-X-1963, W. Teare ( MGCL) . SUDAN: Didinga Mts., Nagichot Sta. 4°15N, 33°39E: 2♀, 6700 ft [2042 m], 16-XII-1925, G. D. H. Carpenter ( MGCL) ; 1♀, 20-XII- 1925, G. D. H. Carpenter ( MGCL) . TANZANIA: Moshi , 1♂, III-[1916], F. C. Selons ( BMNH) . Slopes of Kilamanjaro , 1♂ ( BMNH) . Tanganyika, W. Short of L. Manyara, 1♂, II-V-1935, B. Cooper ( BMNH) . Udzungwa National Park , 1♂, 2-XII-1995, D. Grimaldi ( AMNH) . Usambara , 4♂ ( BMNH) . ZIMBABWE: South Rhodesia: Salisbury : 1♂, 2-IV-1962, A. J. Duke ( AYK) . Umtali : 1♂, 27-I-1948 ( CMNH) . Uncertain country: 1♂ ( CMNH) ; 1♂, 6-III-1945 ( CMNH) . Kalav : 2♂, 25-II-1891 ( CMNH) . Uadoni Park : 1♂, 18-II- 1976 D. M. Kroon ( MGCL) .

Distribution. Eastern and Southern Africa, including Kenya, Malawi, Mozambique, South Africa, Sudan, Tanzania, and Zimbabwe.

Figures

FIGURES 2−13. Lateral view of the male eighth abdominal tergum of Libythea and Libytheana with scales and hairs removed. 2. Libythea celtis; 3. Libythea collenettei; 4. Libythea geoffroyi; 5. Libythea labdaca; 6. Libythea laius; 7. Libythea lepita; 8. Libythea myrrha; 9. Libythea narina; 10. Libytheana carinenta; 11. Libytheana fulvescens; 12. Libytheana motya; 13. Libytheana terena.

FIGURES 27−32. Lateral view of male genitalia of Libythea. 27. Libythea collenettei; 28. Libythea geoffroyi; 29. Libythea labdaca; 30. Libythea laius; 31. Libythea lepita; 32. Libythea myrrha.

FIGURES 101−116. Right dorsal and ventral wing surfaces of Libythea. The second image of each specimen shows the ventral surface of the wings. 101, 102. Libythea geoffroyi howarthi Petersen: Paratype ♂, Solomon Islands, Rennell (BMNH); 103, 104. Libythea labdaca ancoata Grose−Smith: Holotype ♂, Madagascar, Irast (BMNH); 105, 106. Libythea labdaca labdaca Westwood: ♂, Uganda, Mulange (CMNH); 107, 108. Libythea labdaca labdaca Westwood: ♀, Cameroon, Bafia (CMNH); 109, 110. Libythea laius laius Trimen: ♂, Malawi, Cholo (CMNH); 111, 112. Libythea laius laius Trimen: ♀, Malawi, Cholo (CMNH); 113, 114. Libythea laius tsiandava Grose−Smith: Holotype ♂, Madagascar (BMNH); 115, 116. Libythea laius tsiandava Grose−Smith: ♀, Madagascar, Mahabo (BMNH).