Pantoclis globosa, Brazidec & Vilhelmsen, 2022

Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, European Journal of Taxonomy 813, pp. 57-86 : 68-69

Pantoclis globosa sp. nov.

urn:lsid:zoobank.org:act:4BF21BF6-7925-4F2A-8A84-5D8C49BBD8D7

Figs 1D , 3D–H , Table 1

Diagnosis

Eye glabrous; female antenna with: scape 5.5 times as long as wide, pedicel shorter than F1, F1 longest, F2–F13 as long as wide ( Fig. 3D–E ); male antenna with: scape 5.5 times as long as wide, pedicel rounded, F1 longest, emarginate on anterior third, F2–F12 subequal in length ( Fig. 3G–H ); mesosoma sparsely pubescent, shorter than metasoma; pronotum not elongate with epomia extending from pronotal shoulders to front coxa; notauli on mesoscutum complete and convergent posteriorly; propodeum with plicae; marginal vein shorter than radial cell and as long as three quarters of its distance to basal vein; radial cell closed; Rs inconspicuous proximally to r-rs ( Fig. 1D ); petiole as long as wide; gaster globous ( Fig. 3F–G ).

Etymology

The species name refers to the globular gaster of the species. The epithet is to be treated as an adjective.

Type material

Holotype NHMD-608414 , a complete female but partially hidden by a milky coat; paratype NHMD- 608394, a complete male.

Locality and horizon

Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma.

Description

Female

BODY. BL = 2.83 mm. Head oval, subtriangular in frontal view (HeL = 0.48 mm), bearing several short hairs, stouter on frons; eye oval, glabrous; antenna inserted on frontal shelf, at level with lower margin of eyes; toruli well separated; antenna bearing numerous short sensilla; scape shorter than head length, 5.5 times as long as wide; pedicel shorter than F1, longer than wide; 13 flagellomeres; F1 longest, two times as long as wide; F2–F13 as long as wide; F13 tapering at apex (antennomeres length of female holotype, in mm: Sc-0.44; P-0.14; F1-0.16; F2-0.10; F3-0.10; F4-0.09; F5-0.09; F6-0.09; F7-0.09; F8- 0.08; F9-0.09; F10-0.09; F11- 0.09; F12-0.09; F13-0.10); mandibles of ordinary form; occipital carina foveate.

MESOSOMA. Smooth and shiny, shorter than metasoma (MsL = 0.97 mm), sparsely pubescent; pronotum not elongate with distinct and uninterrupted epomia; mesoscutum distinctly convex, notauli complete and convergent; anterior scutellar pit suboval, slightly narrower than apical distance between notauli; scutellum without posterior scutellar pit; propodeum with two prominent plicae; median propodeal keel simple. Fore wing hyaline, homogeneously micropubescent, extending beyond metasoma (FwL = 2.71 mm); C, Sc+R, M+Cu, basal vein, marginal vein, postmarginal vein and r-rs pigmented; Cu nebulous; marginal vein length around three quarters of its distance from basal vein; marginal vein shorter than radial cell; radial cell closed by Rs; Rs pigmented distally to r-rs, inconspicuous proximally. Legs slender, with short hairs; tibial spur formula 1-2-2; tarsal claws simple.

METASOMA. Petiole twice as long as wide (PtL = 0.27 mm; PtW = 0.12 mm), longitudinally striated, bearing long hairs dorsally; gaster globular, slightly longer than high (GL = 1.05 mm; GH = 0.67 mm), with sparse pubescence; S2 and T2 longest; four ring-like segments discernible beyond large tergite.

Male

BL = 3.50 mm; similar to female but antennae sexually dimorphic: scape shorter than head length (HeL = 0.53 mm), 5.5 times as long as wide, pedicel rounded, as long as wide, 12 flagellomeres elongate, cylindrical, longer than wide, F1 longest, with marked emargination on anterior third of segment, F2–F12 subequal in length, F12 tapering at apex (antennomeres length of male paratype, in mm: Sc- 0.38 mm; P-0.11; F1-0.25; F2-0.19; F3-0.19; F4-0.19; F5-0.18; F6-0.17; F7-0.16; F8-0.16; F9-0.16; F10-0.16; F11- 0.15; F12; 0.21).

Remaining measurements: HeL = 0.53 mm; MsL = 1.24 mm; FwL = 2.84 mm; PtL = 0.32 mm; PtW = 0.15 mm; GL = 1.41 mm; GH = 0.82 mm.

Comments

Using Nixon’s (1957) key, Pantoclis globosa sp. nov. keys out to Pantoclis because of the following characters: antennae inserted on a frontal prominence, mandibles not forming a beak or sickle-shaped, notauli present, scutellum without foveae along posterior margin, marginal vein nearly as long as its distance from basal vein and presence of ring-like segments beyond the large tergite, marginal vein shorter than radial cell, epomia distinct, median propodeal keel simple, petiole at most one and a third times as long as its apical width. Pantoclis globosa also fits in Pantoclis according to the diagnosis of Hou et al. (2016). It differs from most extant representatives of Pantoclis by having the marginal vein much longer than the r-rs ( Nixon 1957; Sharma 1980; Buhl 1998). Pantoclis longiscapa Chambers, 1974 has a similar fore wing configuration but differs by having F1 very long and F2 distinctly longer than the following flagellomeres ( Chambers 1974). Pantoclis deperdita Brues, 1906 , from the Eocene of Florissant has a shorter radial cell and marginal vein ( Brues 1906) and is therefore considered as a different species from P. globosa .

Figures

Fig. 1. Wing venation of the species described here. A. Belyta knudhoejgaardi sp. nov. B. Cinetus breviscapus sp. nov.C. Cinetus elongatus sp. nov.D. Pantoclis globosasp. nov. E. Pantolyta augustinusii sp. nov. F. Basalys villumi sp. nov. G. Doliopria baltica sp. nov. H. Spilomicrus succinalis sp. nov. Scale bars: A–E = 1 mm; F, H = 0.5 mm; G = 0.25 mm.

Fig. 3. A–C. Cinetus elongatus sp. nov., holotype (NHMD-608402). A. Habitus in lateral view. B. Head in lateral view. C. Mesosoma and petiole in lateral view. D–H. Pantoclis globosa sp. nov. D–F. Holotype (NHMD-608414). D. Habitus in lateral view. E. Habitus in dorsal view. F. Gaster in lateral view. G–H. Paratype (NHMD-608394). G. Habitus in lateral view. H. Habitus in dorsal view. Abbreviations: asp = anterior scutellar pit; fe = flagellomere emargination; mtr = metapleural ridge. Scale bars: A = 2 mm; B–E, G–H = 1 mm; F = 0.5 mm.