Panscopus (Phymatinus) spantoni Bright, 2008

Spanton, Timothy G. & Anderson, Robert S., 2016, A Taxonomic Revision of Weevils of the Genus Panscopus Schönherr (Coleoptera: Curculionidae: Entiminae: Tropiphorini), The Coleopterists Bulletin (mo 15) 70, pp. 1-86 : 32-35

Panscopus (Phymatinus) spantoni Bright, 2008

(Frontispiece, Plate 2 , Figs. 11 , 37 , 60 , Map 12 )

Panscopus (Nocheles) torpidus ; Pierce 1913: 390 (not LeConte: 1857); Leng 1920: 312; Buchanan 1927: 28, 1936: 9; Schenkling and Marshall 1931: 40; Ting 1938: 122, 123; Blackwelder 1939: 66; Bousquet et al. 2013: 345.

Panscopus torpidus ; Hatch 1971: 289; O’ Brien and Wibmer 1982: 63.

Panscopus spantoni Bright (in Bright and Bouchard 2008: 264).

Type Material. Holotype female ( CASC), allotype male ( CASC), and 26 paratypes ( CASC, CNCI) bear the label data: “Olympia/Wash. VII-28-28; Pres by/Thos Craig/ Collector”.

Notes about Synonymy. The holotype of Hylobius torpidus LeConte, 1857 is a specimen of the species that has been recognized as Panscopus pallidus Buchanan, 1927 by Buchanan (1936), Hatch (1971), and O’ Brien and Wibmer (1982). Thus, the name Panscopus pallidus Buchanan is a junior synonym of Panscopus (Nomidus) torpidus (LeConte) . See the discussion under that species name.

Specific Epithet. The species name is a latinized genitive patronym of the surname of the first author, Timothy Gordon Spanton, recognizing his contribution in advancing understanding of the classification and identification of the genus Panscopus .

Diagnosis. Most specimens of this species are recognized by the following combination of character states: scape long, extending to hind margin of eye; four ventrites in abdomen of females; alternate elytral intervals distinctly costate, even intervals nearly non-setose; elytral setae thin, but flattened; color black to cinereous. In addition, males exhibit in the internal sac a field of spicules distad belt of tegmen and a v-shaped pair of sclerites ( Fig. 60 ).

Redescription. Females: Length 8.1–9.9 mm (n = 10), width across elytra 3.4–4.1 mm (n = 10). Males: Length 7.7–9.4 mm (n = 10), width across elytra 3.0– 3.6 mm (n = 10). Color: Very dark brown to black, with areas of medium or light brown scales posteriorly and laterally on elytra. Head: Transverse concavity at base of frons nearly absent; rostrum and frons apparently continuous in lateral view; rostrum with thin, shiny longitudinal median carina and lateral sulci indistinct dorsally in apical portion, in most specimens; nasal plate not clearly defined from punctate surface of rostrum, v-shaped carina on nasal plate; vestiture of frons and rostrum of scattered decumbent to appressed setae. Antennae: Scape as long as or longer than rostrum, extending to or beyond hind margin of eye, vestiture of apically directed reclinate setae, integument visible among the setae; antennomeres 2–8 with long primary setae on distal half of each article and scattered, shorter, finer, setae; shiny integument clearly visible through thin vestiture. Pronotum: Slightly wider than long, roundly convex laterally, widest slightly anterior of middle; surface coarsely rugose-tuberculate; median longitudinal sulcus nearly length of pronotum; vestiture dorsally of scattered, decumbent to appressed setae located at apices of low rounded tubercles, scales absent, or nearly so, dorsally; laterally some scattered, appressed, striate scales present. Elytra: Elongate oval, combined width 1/3–1/4 wider than pronotum; declivity, in lateral view, nearly vertical in males, reflexed in females; alternate intervals raised, with irregular, double to partly triple row of decumbent, thin setae, serial pits each with scale only slightly larger than surrounding scales; elsewhere, vestiture of dense, overlapping, appressed, striate scales. Legs: Tibiae 1 and 2 with small denticles on ventral margin in distal half, setae emanating therefrom not markedly stiffened and spine-like, in most specimens; tibia 3 without spines on ventral margin apically, corbellar area indistinctly closed at apex of tibia 3. Abdomen: With 4 ventrites in females (ventrites 4 and 5 fused); 5 ventrites in males; in females, ventrite 1 convex at middle; in males, concave at middle. Genitalia: Females ( Figs. 11 , 37 ): Sternum VIII (spiculum ventrale) with apical expansion broadly rounded apically, convex ventrally and wide and shovel-shaped, cleft medially, with thin setae near apex; coxites 1 and 2 broadly angular, narrowed apically in dorsal and ventral views, rounded apically in lateral view; styli each with 2 or 3 primary setae at apex; vagina shorter than length of coxites, membranous, with pair of broadly crescent-shaped sclerites near confluence of common oviduct; spermatheca sickle-shaped, thick over most of its length. Males ( Fig. 60 ): Aedeagus with median lobe greatly decurved, convex dorsally, concave ventrally in retracted state; apex with broad, flat, spatulate projection, in dorsal view, narrow and acute in lateral view; manubrium of tegmen approximately as long as median lobe; median struts of aedeagus slightly longer than median lobe; internal sac membranous with small pair of sclerotized structures near apex of sac (anteriorly in inverted sac).

Life History. Label data indicate that most adults have been found from March through October. Isolated records are from the months of February, November, and December, the latter two records from litter.

Plant Associations. Records were seen of specimens being taken from alder, dandelion, red clover, strawberry, as well as litter of hemlock, oak, and vine maple, and on moss near logs.

Geographical Distribution. This species lives in the southern mainland of British Columbia (Vancouver) and in the moist forest regions of western Oregon and Washington ( Map 12 ). Of the few elevation records available, the highest are 950–1,070 m .

Chorological Affinities. The ranges of this species, P. squamosus , and P. michelbacheri are adjacent to one another but non-overlapping.

Material Examined. In addition to the type specimens listed above, approximately 240 specimens were examined. CANADA: British Columbia: “B.C. V.” ( Vancouver ), 4.VII.1937, 1 ( OSAC) . USA: Oregon: Benton Co., “Campus” ( OSAC campus, Corvallis), (1, OSAC) ; Corvallis , 12.VI.1933, J. Schuh (2, AMNH) ; McDonald Forest, N. of Corvallis , 3.XI.1949, V. Roth (1, OSAC) . Clatsop Co., Birkenfeld , 8 km S.W., 30.X.1960, R. H. Heitmanek, rotten wood & ground litter (6, OSAC) ; Cannon Beach , 15.VI.1927, E. C. Van Dyke (1, CASC) ; J.J. Astor Experiment. Station , 29.IX.1960, E. Dickason, moss on logs (1, OSAC) . Columbia Co., Goble , 18.V.1938, Gray & Schuh (2, AMNH) ; Goble , 22.IV.1938, K. Gray & J. Schuh, (Emergence cages) (1, USNM) . Douglas Co., Reedsport , 20.VI.1939, T. G. Aitken (1, AMNH) . Jackson Co., Union Creek 950–1070 m, 1-15.IX.1950, B. Malkin (1, FMNH) . Josephine Co., Silver Creek Falls , 1.VIII.1948, K. M. Fender (1, OSAC) . Lane Co., Eugene , 24.VII.1941, 8.IX.1941, VII.1941, B. Malkin (4, FMNH) ; Junction City , 16 km N.W., IX.1947, R. O. Mattus (1, FMNH) ; Spencer’ s Butte, Eugene , VII.1942, B. Malkin (2, FMNH) . Linn Co., Albany , 10.VIII.1929, J. Wilcox, Strawberry (1, OSAC) ; Lacombe , 3 mi E., 13.XII.1957, Quercus litter (1, OSAC) . Marion Co., Mill Valley , 18.VI.1925, E. P. Van Duzee (1, CASC) ; Silver

Falls State Park , 23.VI.1939, M.A. Cazier collection (3, AMNH) . Multnomah Co., Gresham , 18. IV.1946, 7.IV.1947, Joe Schuh, Strawberry (2, AMNH) ; Portland, 22&23.V, Hubbard & Schwarz (4, USNM) . Washington Co., Garden Home , 23. V.1919, L. P. Rockwood (1, USNM) ; North Plains , 23.IV.1959, K. Goeden, forest litter (1, ODAC) ; Orenco , 4.VI.1945, Anderson, Alnus (2, USNM) ; Pumpkin Ridge, 24.IV.1973, Westcott & Penrose, Red Clover (2, CWOB; 1, ODAC). Yamhill Co., Dayton , 3.VI.1942, 12.IV.1949, K. M. Fender (2, OSAC) ; County unknown: “Or” (3, CMNH) . Washington: Clallam Co., Forks , 2.VII.1920, Van Dyke collection (1, CASC) ; Klayhowha Campground, Olympic National Forest , 16.VIII.1988, R. Baranowski (1, CMNC) . Grays Harbor Co., Humptulips , 29.V.1914, Van Dyke collection (1, CASC) ; Montesano, 20.III.1932, J. Wilcox (2, UNSM; 10 OSAC); Montesano, 3.IV.1932, J. Wilcox (2, CNCI; 26, OSAC; 1, UCDC); Montesano , 8.V.1932, J. Wilcox, 4 (4, OSAC; 5, TAMU; (1, UCDC) ; Montesano , 25.II.1933, J. Wilcox (5, OSAC) ; Montesano , 14.V.1933, J. Wilcox (11, OSAC) . King Co., Seattle , 9.V.1907, Van Dyke collection (5, CASC) ; 10.V.1953, 16.V.1953, B. Malkin (2, FMNH) ; Seattle , 21.IV.1938, 9.V.1938, 25.V.1938, E. Dailey (4, OSAC) ; Seattle , 14.IV. 1938, C. H. Lovers (1, OSAC) ; Seattle , 26.VI.1929, M. H. Hatch (1, OSAC) ; Seattle , 23.IV.1939, J. Stort (1, OSAC) ; Seattle , 9.IV.1924, 6.IV.1925, 17.V.1933, 2.VI.1933 (6, OSAC) . Lewis Co., Lewis and Clark State Park , 5.IX.1959, K. M. Fender (1, OSAC) . Pierce Co., Puyallup , 8.V.1939, S. Crumb (1, OSAC) ; Puyallup , 30.IV.1930, W. W. Baker (5, USNM) ; Puyallup , 21.V.1931, W.W. Baker (1, WIRC) ; Summer , 23.IV.1953 (8, USNM) ; Tacoma , 25.IV.1931, W. W. Baker (1, CUIC) . Thurston Co., East Olympia , 11.V.1945, Forsell, Dandelion (1, USNM) ; Olympia, 16.III, 10.IV, 26. VI, Liebeck collection (1, MCZC) ; Olympia, Wickham collection (4, USNM) ; Olympia , 4.IX. 1925, M. C. Lane (1, USNM) ; Rainier , 24.IV. 1914, W. J. Chamberlain (1, USNM) ; Rochester , 3.IV.1931, 30.III.1932, W. W. Baker (2, CNCI) ; Rochester, 19.IV.1930, W. W. Baker (3, CUIC; 2, UCDC; 7, USNM; 1, WSU); Rochester , 19.IV. 1930, W. W. Baker (1, CUIC) ; Rochester , 30.IV. 1930, W.W. Baker (1, WIRC) ; Rochester , 3.V.1930, W. W. Baker (2, USNM) . County unknown: “WA”, Van Dyke collection (1, CASC) ; “WA”, F. R. Eddy collection (1, MCZC) ; “W. Ter. ”, 20.III.1892 (1, USNM) ; “W.Ter.” (1, USNM) .

Figures

Plate 2. Panscopus spantoni, dorsal habitus. From Bright and Bouchard (2008), used with permission.

Map 12. Geographical distribution of Panscopus spantoni.

Figs. 9–16. Panscopus species, sternum 8 (spiculum ventrale), ventral (a) and left lateral (b) views. 9) P. acutisetus; 10) P. gemmatus; 11) P. spantoni; 12) P. squamosus; 13) P. michelbacheri; 14) P. johnsoni; 15) P. bakeri; 16) P. longus. Scale bar = 1 mm.

Figs. 37–40. Panscopus species, coxites and vagina, dorsal (a), ventral (b), and left lateral (c) views. 37) P. spantoni;

Figs. 59–61. Panscopus species, aedeagus, dorsal (a), ventral (b), and left lateral (c) views. 59) P. gemmatus; 60) P. spantoni; 61) P. squamosus. b = belt of tegmen; is = internal sac; mb = manubrium of tegmen; ml = median lobe; ms = median strut; t = tegmen. Scale bar = 1 mm.