Lanceobostra, Hennemann & Conle, 2024

Hennemann, Frank H. & Conle, Oskar V., 2024, Studies on Neotropical Phasmatodea XXVI: Taxonomic review of Cladomorformia tax. n., a lineage of Diapheromerinae stick insects, with the descriptions of seven new genera and 41 new species (Phasmatodea: Occidophasmata: Diapheromerinae), Zootaxa 5444 (1), pp. 1-454 : 98-99

19. Lanceobostra (?) simplex ( Redtenbacher, 1908: 418) [ Bacteria ]. HT, ♀: Mexiko [SMNS—not traced]. comb. n. Remarks: Redtenbacher (1908: 418) estimated this species might be a synonym of L (?) foliolata ( Redtenbacher, 1908) . Distribution: Mexico [SMNS—not traced].

Keys to the species of Lanceobostra gen. n.

♀♀ *

1. Head sub-cylindrical.................................................................................. 2

- Head ovoid with vertex weakly convex ( Fig. 38H ); Oaxaca ......................................... oaxacae sp. n.

2. Small insects (body length incl. subgenital plate <140.0 mm................................................... 3

- Larger (body length incl. subgenital plate> 150.0 mm)........................................................7

3. Meso- and metanotum densely tubercular or spinulose........................................................ 4

- Meso- and metanotum only with a few scattered tubercles or granules............................................ 6

4. Meso- and metasternum granulose or tubercular; head and mesofemora unarmed; Jalisco ............................ 5

- Meso- and metasternum spinulose; head with a large pair of auriform lobes; mesofemora with two small rounded sub-basal lobes on two outer ventral carinae; Colima ........................................................... frustrans

5. Body surface dull; meso- and metanotum densely and evenly granulose; mesonotum with a longitudinal, granulose lateral carina........................................................................................... similis

- Body surface glossy ( Figs. 38A–B ); meso- and metanotum unevenly tubercular ( Fig. 38G ); no longitudinal lateral carina along mesonotum................................................................................. glabra sp. n.

6. Green; body surface glossy; head 1.3x longer than wide; medioventral carina of meso- and metafemora with three distinct teeth; Guerrero .................................................................................. tridenticulata

- Brown; body surface dull; head elongate, compressed dorsoventrally and 1.5x longer than wide; medioventral arina of meso- and metafemora with 2–3 minute denticles; Cuba (?).................................................. .. deplanata

7. Head unarmed or with two cephalad spines or auriform lobes.................................................. 8

- Head with a pair of acute conical cephalad horns and an additional pair of slender, bifid spines between the eyes; Michoacan ...................................................................................... chapalaense sp. n.

8. Lateral margins of abdominal tergum VII with a posterolateral lobe.............................................. 9

- Abdominal tergum VII with lateral margins straight......................................................... 10

9. Meso- and metanotum unevenly tubercular; meso- and metatibiae with a two large almost semi-circular dorsal lobes ( Figs. 36D–E ); dorsal lobe of basitarsi prominent and roundly triangular; Oaxaca ............................... ornata sp. n.

- Meso- and metanotum rather evenly granulose; meso- and metatibiae at best with a small dorsal tooth or Lobe about one-third off the base; dorsal lobe of basitarsi small, and rounded; Sinaloa & Sonora ................................... aetolus

10. Head <1.3x longer than wide and bi-cornute; Guatemala..................................................... 11

- Head elongate,> 1.5x longer than wide and unarmed; Mexico (Nayarit ................................... margaritata

11. Very slender species, mesonotum 8.2x longer than pronotum; cephalad horns with apex bifid; basitrarsi Crested dorsally............................................................................................... freygessneri

- Stockier species, mesonotum only 7.2x longer than pronotum; cephalad horns conical; basitarsi simple........ guatemalensis

[* ♀♀ of L. jaliscensis ( Rehn, 1904) comb. n., and L. mirata ( Redtenbacher, 1908) comb. n., L. torqata sp. n. and L. tuckerae sp. n. are not known]

♂♂ *

1. Very large (body length 160.0 mm); Guatemala.......................................................... mirata

- Smaller.............................................................................................. 2

2. Cerci round in cross-section and ± club-shaped.............................................................. 3

- Cerci distinctly shovel- or paddle-shaped and canaliculate interiorly............................................. 4

3. Head ovoid ( Fig. 38J ); lateral margins of abdominal tergum IX straight ( Fig. 39R ); SW-Mexico (Oaxaca)..... oaxacae sp. n.

- Head sub-cylindrical; lateral margins of abdominal tergum IX rounded ( Fig. 40Q ); Guatemala.............. guatemalensis

4. Cerci straight along the longitudinal axis; posterior margin of anal segment not truncated.............................5

- Cerci torquate and twisted along longitudinal axis ( Figs. 39 O-Q); posterior margin of anal segment truncated ( Fig. 39P )............................................................................................. torquata sp. n.

5. Poculum with a prominent central spiniform protuberance..................................................... 6

- Poculum with central protuberance weakly developed and obtuse ( Fig. 40D ); W-Mexico (Michoacan).... chapalaense sp. n.

6. Poculum projecting notably beyond posterior margin of abdominal tergum IX; anal segment with posterior margin notably narrowed............................................................................................ 7

- Poculum at best reaching to posterior margin of anal segment; anal segment not narrowed............................ posteriorly............................................................................................8

7. Anal segment with one ventral thorn-pad with entire posteroventral surface denticulate; W-Mexico..................... (Najarit).................................................................................... margaritata

- Anal segment with two ventral thorn-pads with only outer lateral portions of posterior margin denticulate Ventrally ( Fig. 95C ); NW-Mexico (Sinaloa & Sonora)...................................................................... aetolus

8. Posterior margin of anal segment rounded ( Fig. 39V )............................................. .. tuckerae sp. n.

- Anal segment different................................................................................. 9

9. Posterior margin of anal segment tri-lobate with the central portion protruded ( Fig. 40O )......................... similis

- Posterior margin of anal segment with a wide concave excavation ( Fig. 40H )............................... jaliscensis

[* ♂♂ of L. deplanata ( Redtenbacher, 1908) comb. n., L. freygessneri ( Redtenbacher, 1908) comb. n., L. frustrans ( Redtenbacher, 1908) comb. n., L. glabra sp. n., D. ornata sp. n. and L. tridenticulata ( Redtenbacher, 1908) comb. n. are not known]

Figures

FIGURE 36. Lanceobostra gen. n.: A. L. aetolus (Westwood, 1859) ♀ with lobed legs (Mexico, Sinaloa, Venvidio), habitus dorsolateral view [ANSP]; B. L.aetolus (Westwood, 1859) ♀ with unarmedlegs (Mexico,Sinaloa,Venvidio),habitus dorsolateral view [ANSP]; C. L. aetolus (Westwood, 1859) ♂ (Mexico. Sinaloa, Venvidio), habitus lateral view [ANSP]; D. L. ornata sp. n. ♀ PT, habitus dorsolateral view [USNM, © Paul D. Brock]; E. L. ornata sp. n. ♀ PT, habitus lateral view [USNM, © Paul D. Brock]; F. L. ornata sp. n. ♂ HT, habitus dorsolateral view [ANSP]; G. L. guatemalensis (Bruner v. Wattenwyl, 1907) ♂, habitus dorsolateral view [ANSP].

FIGURE 38. Lanceobostra gen. n.: A. L. glabra sp. n. ♀ HT, habitus dorsal view [UMMZ, © Erika Tucker]; B. L. glabra sp. n. ♀ HT, habitus dorsal view [UMMZ, © Erika Tucker]; C. L. torquata sp. n. ♂ HT, habitus dorsolateral view [ZMUH]; D. L. aetolus (Westwood, 1859) ♀, head with small tubercles in dorsolateral view [ANSP]; E. L. aetolus (Westwood, 1859) ♀, head with prominent multi-dentate cephalic horns in dorsolateral view [ANSP]; F. L. glabra sp. n. ♀ HT, head and prothorax in lateral view [UMMZ, © Erika Tucker]; G. L. glabra sp. n. ♀ HT, head, pronotum and anterior half of mesonotum in dorsal view [UMMZ, © Erika Tucker]; H. L. oaxacae sp. n. ♀ HT, head and prothorax in dorsolateral view [ANSP]; J. L. oaxacae sp. n. ♂ PT, head and prothorax in dorsolateral view [coll. FH]; K. L. aetolus (Westwood, 1859) ♂, head in lateral view [ANSP]; L. L. chapalaense sp. n. ♂ PT, head and prothorax in dorsolateral view [ANSP]; M. L. tuckerae sp. n. ♂ PT, head and prothorax in lateral view [UMMZ, © Erika Tucker]; N. L. torquata sp. n. ♂ HT, head and prothorax in dorsal view [ZMUH]; O. L. torquata sp. n. ♂ HT, head and prothorax in lateral view [ZMUH].

FIGURE 39. Lanceobostra gen. n. terminalia: A. L. aetolus (Westwood, 1859) ♀, terminalia in lateral view [ANSP]; B. L. aetolus (Westwood, 1859) ♀, terminalia in dorsal view [ANSP]; C. L. aetolus (Westwood, 1859) ♀ with lobed abdominal tergum VII, terminalia in lateral view [ANSP]; D. L. aetolus (Westwood, 1859) ♀ with lobed abdominal tergum VII, terminalia in ventral view [ANSP]; E. L. aetolus (Westwood, 1859) ♀, terminalia in lateral view [ANSP]; B. L. glabra sp. n. ♀ HT, terminalia in lateral view [UMMZ, © Erika Tucker]; F. L. glabra sp. n. ♀ HT, terminalia in dorsal view [UMMZ, © Erika Tucker]; G. L. chapalaense sp. n. ♀ HT, terminalia in lateral view (tip of subgenital plate broken off) [ANSP]; H. L. chapalaense sp. n. ♀ HT, terminalia in dorsal view (tip of subgenital plate broken off) [ANSP]; J. L. chapalaense sp. n. ♀ HT, terminalia in ventral view (tip of subgenital plate broken off) [ANSP]; K. L. guatemalensis (Brunner v. Wattenwyl, 1907) ♀ HT of Bostra intermedia Redtenbacher, 1908, terminalia in lateral view [MHNG]; L. L. oaxacae sp. n. ♀ HT, terminalia in lateral view [ANSP]; M. L. oaxacae sp. n. ♀ HT, terminalia in dorsal view [ANSP]; N. L. oaxacae sp. n. ♀ HT, terminalia in ventral view [ANSP]; O. L. torquata sp. n. ♂ HT, terminalia in lateral view [ZMUH]; P. L. torquata sp. n. ♂ HT, terminalia in dorsal view [ZMUH]; Q. L. torquata sp. n. ♂ HT, terminalia in ventral view [ZMUH]; R. L. oaxacae sp. n. ♂ PT, terminalia in lateral view [coll. FH]; S. L. oaxacae sp. n. ♂ PT, terminalia in dorsal view [coll. FH]; T. L. oaxacae sp. n. ♂ PT, terminalia in ventral view [coll. FH]; U. L. tuckerae sp. n. ♂ PT, terminalia in lateral view [UMMZ, © Erika Tucker]; V. L. tuckerae sp. n. ♂ PT, terminalia in dorsal view [UMMZ, © Erika Tucker].

FIGURE 40. Lanceobostra gen. n. terminalia of ♂♂: A. L. aetolus (Westwood, 1859), lateral view [ANSP]; B. L. aetolus (Westwood, 1859), dorsal view [ANSP9; C. L. aetolus (Westwood, 1859), ventral view [ANSP]; D. L. chapalaense sp. n. ♂ PT, lateral view [ANSP]; E. L. chapalaense sp. n. ♂ PT, dorsal view [ANSP]; F. L. chapalaense sp. n. ♂ PT, ventral view [ANSP]; G. L. jaliscensis (Rehn, 1904), lateral view [ANSP]; H. L. jaliscensis (Rehn, 1904), dorsal view [ANSP]; J. L. jaliscensis (Rehn, 1904), ventral view [ANSP]; K. L. ornata sp. n. ♂ HT, lateral view [ANSP]; L. L. ornata sp. n. ♂ HT, dorsal view [ANSP]; M. L. ornata sp. n. ♂ HT, ventral view [ANSP]; N. L. similis (Redtenbacher, 1908), lateral view [ANSP]; O. L. similis (Redtenbacher, 1908), dorsal view [ANSP]; P. L. similis (Redtenbacher, 1908), ventral view [ANSP9; Q. L. guatemalensis (Brunner v. Wattenwyl, 1907), lateral view [ANSP]; R. L. guatemalensis (Brunner v. Wattenwyl, 1907), dorsal view [ANSP]; S. L. guatemalensis (Brunner v. Wattenwyl, 1907), ventral view [ANSP].

FIGURE 95. Vomer of ♂♂ (ventral view): A. Calynda coronata Carl, 1913 [ANSP]; B. Spinocloidea panamaense gen. n., sp. n. PT [coll. FH]; C. Lanceobostra aetolus (Westwood, 1859) [ANSP]; D. Lanceobostra ornata sp. n. HT [ANSP]; E. Lanceobostra jaliscensis Rehn, 1904 [ANSP]; F. Lanceobostra guatemalensis (Redtenbacher, 1908) [ANSP]; G. Bostriana boliviana (Piza, 1939) [coll. OC]; H. Alienobostra remiformis (Rehn, 1904) LT of Bostra jugalis (Rehn, 1905) [ANSP]; J. Phanoclocrania speciosa Conle, Hennemann & Gutiérrez, 2011 [CEBUC, © Yeisson Gutierréz]; K. Phanoclocrania dorsuaria (Stål, 1875) [ZMUH]; L. Parotocrania panamae gen. n., sp. n. PT [coll. OC]; M. Parotocrania regina (Conle, Hennemann & Gutiérrez, 2011 [CEBUC, © Yeisson Gutierréz]; N. Globocrania emesa (Westwood, 1859) HT of Pseudobacteria longipes Piza, 1938 [MZUSP]; O. Globocrania culmus (Bates, 1865) [CEIOC]. Not in scale.