Phanocles mutica (Brunner von Wattenwyl, 1907)

Hennemann, Frank H. & Conle, Oskar V., 2024, Studies on Neotropical Phasmatodea XXVI: Taxonomic review of Cladomorformia tax. n., a lineage of Diapheromerinae stick insects, with the descriptions of seven new genera and 41 new species (Phasmatodea: Occidophasmata: Diapheromerinae), Zootaxa 5444 (1), pp. 1-454 : 149-150

20. Phanocles mutica (Brunner v. Wattenwyl, 1907: 324 ) [ Dyme ]. HT, ♂: Coll. Br. v. W., Trinidad, Boucard; det. Br. v. W. Dyme mutica ; 12.852 [NHMW, No. 652]. ( Figs. 53C , 55G , 57E , 60C , 64 H-K, 66M-O, 92F, 97N, 99F, 118D–F)

= Bacteria bradypus Redtenbacher, 1908: 421 . LT [by present designation], ♀: Museum Paris, Antilles; det. Redtenb. Bacteria bradypus ; 23.325 [NHMW, No. 818]. syn. n. (Not: PLT, ♀: det. Redtenb. B. Bellangeri ; B. bicornis Stoll. — Burm. Surin. [NHMW, No. 818] → This is Phanocles maroniensis ( Chopard, 1911))

= Bacteria integra Redtenbacher, 1908: 421 . HT, ♀: Coll. Br. v. W., Trinidad, Boucard; det. Redtenb., B. integer ; 12.853 [NHMW, No. 811].

Remarks: Differs from keratosqueleton by the slenderer body and relatively longer body segments and slightly longer median segment of both sexes (> 2x length of metanotum in ♂♂); large triangular dorsal lobe of the basitarsi of ♀♀, as well as the unarmed head and anal segment of ♂♂ which is shorter and not dilated towards the posterior. Bacteria integra Redtenbacher, 1908 has erroneously been synonymised with P. keratosqueleton ( Olivier, 1792) by Zompro (1998: 6). Langlois & Bellanger (2012: 103) recorded Ph. muticus from Tobago as “ Paraphanocles keratosqueleton ”.

Distribution: Trinidad: Trinidad [MNHN, MNHU, NHMW]; Trinidad, Anima Valley, Simla, W Beebe Tropical Research Station [FSCA]; Trinidad, Fyzabad [ANSP]; Trinidad, Caparo [ANSP]; Trinidad, Mount Tunapuna, St. Benedict [ANSP]. Tobago: Tobago, trace à 3, 2 km á l’ouest de Castara, route de Plymouth [MNHN, coll. ASPER]; Tobago, trace au sud du barrage de Hillsborough, prés de Green Hill [MNHN, coll. ASPER]; Tobago, Black Rock [ASPER]; Tobago, route de Branch, près de Masson Hall [ASPER]. “Antilles” [NHMW].

Figures

FIGURE 53. Phanocles Stål, 1875 habitus: A. P. pleurospinosus sp. n. ♀ HT (Costa Rica, Prov. Limón, Pococí, Bosque Lluvioso), lateral view [MNCR-A]; B. P. pleurospinosus sp. n. ♀ HT, dorsal view [MNCR-A]; C. P. muticus (Brunner v. Wattenwyl, 1907) ♀ (captive reared from Trinidad), dorsolateral view [coll. FH, No. 0837–13]; D. P. barbadosense sp. n. ♀ HT (Barbados), dorsal view [NHMW]; E. P. barbadosense sp. n. ♀ HT (Barbados), dorsolateral view [NHMW, © 2019 Natural History Museum Vienna, NOaS Image Collection / H. Bruckner]; F. P. acutecornutus sp. n. ♀ HT (Ecuador, Prov. Napo, Yasuni), dorsal view [MNHN]; G. P. acutecornutus sp. n. ♀ HT (Ecuador, Prov. Napo, Yasuni), dorsolateral view [MNHN].

FIGURE 55. Phanocles Stål, 1875: A. P. ecuadoricus sp. n. ♂ PT (Ecuador, Prov. Esmeraldas, Rio Verde), dorsolateral view [MNCN]; B. P. falcatus sp. n. ♂ PT (Ecuador, Prov. Napo, Misahualli), dorsolateral view [coll. FH, No. 0919–3]; C. P. maximus sp. n. ♀ PT (captive reared from Panamá, Prov. Coclé, Valle deAntón), dorsolateral view [coll. FH, No. 1360–3]; D. P. spectabilis sp. n. ♂ PT (captive reared from Costa Rica, Prov. Puntarenas, La Gamba), dorsolateral view [coll. FH, No. 0563–18]; E. P. ploiaria (Westwood, 1859) ♀ (Panamá, Prov. Coclé, Valle de Antón 650–850 m), dorsolateral view [coll. FH, No. 0705–12]; F. P. obtusus (Redtenbacher, 1908) ♂ (Venezuela, Aragua N.P., Rancho Grande), dorsolateral view [ANSP].

FIGURE 57. Phanocles Stål, 1875 head and prothorax: A. P. acutecornutus sp. n. ♀ HT, dorsolateral view [MNHN]; B. P. berezini sp. n. ♀ PT, lateral view [coll. FH]; C. P. brevipes sp. n. ♀ HT, dorsolateral view (head is turned upside down) [ANSP]; D. P. barbadosense sp. n. ♀ PT, lateral view [ANSP]; E. P. muticus (Brunner v. Wattenwyl, 1907) ♀, dorsolateral view [coll. FH]; F. P. burkartii (Saussure, 1868) ♀, dorsolateral view [coll. FH]; G. P. saussurei (Redtenbacher, 1908) ♀, lateral view [coll. FH]; H. P. falcatus sp. n. ♀ PT, dorsolateral view [MNHN]; J. P. falcatus sp. n. ♀ PT, dorsolateral view [MNHN]; K. P. superbus sp. n. ♀ HT, dorsolateral view [MNHN].

FIGURE 60. Phanocles Stål, 1875 head and prothorax: A. P. aequatorialis (Redtenbacher, 1908) ♂, lateral view [coll. FH]; B. P. costaricensis Hennemann, 2002 ♂, lateral view [coll. FH]; C. P. muticus (Brunner v. Wattenwyl, 1907) ♂, lateral view [coll. FH]; D. P. mexicanus sp. n. ♂ PT, lateral view [UMMZ, © Erika Tucker]; E. P. horni (Redtenbacher, 1908) ♂, lateral view [coll. FH]; F. P. maximus sp. n. ♂ PT, dorsolateral view [coll. FH]; G. P. spectabilis sp. n. ♂ PT, lateral view [coll. FH]; H. P. saussurei (Redtenbacher, 1908) ♂, lateral view [coll. FH]; J. P. vosseleri (Redtenbacher, 1908) ♂, lateral view [coll. FH].

FIGURE 64. Phanocles Stål, 1875 terminalia of ♀♀: A. P. superbus sp. n. ♀ HT, lateral view [MNHN]; B. P. superbus sp. n. ♀ HT, dorsal view [MNHN]; C. P. superbus sp. n. ♀ HT, ventral view [MNHN]; D. P. acutecornutus sp. n. ♀ PT, lateral view [ZSMC]; E. P. berezini sp. n. ♀ PT, lateral view [coll. FH]; F. P. berezini sp. n. ♀ PT, dorsal view [coll. FH]; G. P. berezini sp. n. ♀ PT, ventral view [coll. FH]; H. P. muticus (Brunner v. Wattenwyl, 1907), lateral view [coll. FH]; J. P. muticus (Brunner v. Wattenwyl, 1907), dorsal view [coll. FH]; K. P. muticus (Brunner v. Wattenwyl, 1907), ventral view [coll. FH]; L. P. horni (Redtenbacher, 1908), lateral view [coll. FH]; M. P. horni (Redtenbacher, 1908), dorsal view [coll. FH]; N. P. horni(Redtenbacher, 1908), ventral view [coll. FH]; O. P. mexicanus sp. n. ♀ PT, lateral view [MNHN]; P. P. mexicanus sp. n. ♀ PT, dorsal view [MNHN]; Q. P. mexicanus sp. n. ♀ HT, ventral view [MNHN]; R. P. barbadosense sp. n. ♀ PT, lateral view [ANSP]; S. P. barbadosense sp. n. ♀ PT, ventral view [ANSP].