Fortipalpa sacha, Supeleto & Santos & Aguiar, 2022

Supeleto, Fernanda A., Santos, Bernardo F. & Aguiar, Alexandre P., 2022, Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae), Zootaxa 5219 (6), pp. 501-533 : 521-523

Fortipalpa sacha sp. nov.

( Figs 12–13 , 19–21 )

Description. Holotype ♀. Fore wing 8.65 mm. Head ( Figs 12 , 13A–B ). MLW 1.54. MWW 0.49. MSM 0.25. CHW 0.50. Clypeus anterior margin centrally with two small tubercles. Antenna with 22 flagellomeres. Supra-antennal area distinctly coriarious; median line distinct and straight.

Thorax ( Figs 12 , 13A ). Epomia approximately straight. Mesoscutum length to width 0.97 × as long as wide. Notaulus reaching about 0.54 of mesoscutum length. Scuto-scutellar groove with distinct longitudinal wrinkles. Epicnemial carina reaching about 0.8 of distance to subalar ridge; ending above pronotoal spiracle; slightly curved. Sternaulus moderately strong, wide and shallow at posterior 0.25; strongly sinuous. Mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate-rugulose; moderately pilose.

Propodeum ( Figs 12 , 13A&C ). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elongate, SLW 2.21. Anterior transverse carina medially slightly arched forwards.

Wings ( Fig. 12 ). Fore wing crossvein 1m-cu basally bent, otherwise straight; limit between 1m-cu and 1-Rs+M more or less traceable. Vein 2Cua 0.79 × as long as crossvein 2cu-a. APH 0.56. Areolet subpentagonal; distinctly wider than high, AWH 0.79. Crossvein 3r-m nebulous. Crossvein 2r-m and 3r-m about same length. Vein 3-M distinctly shorter than 2-M. HW1C 1.19. Hind wing vein Cub almost straight, very slightly convex. Vein 2-1A ending at 0.20 of distance to wing margin.

Metasoma ( Figs 12 , 13A&D ). T1LW 2.33. T1WW 2.22. T1 spiracle at 0.54 of its length. Median depression of T1 elongate, very shallow. T2LW 0.98. T2WW 1.91. T2–8 from distinctly coriarious T2 to smooth T8. Ovipositor straight, OST 1.70; ventral valve apex with 15 teeth ( Fig. 13D ).

Color. Head pale yellow (233,215,149) with supra-antennal area and vertex black (36,31,21). Clypeus mid ventrally with dark brown spot. A moderately wide dark brown stripe extending from tentorial pit to supraclypeal area. Orbital band almost complete, briefly interrupted near 6 h. Mandible pale yellow, apical 0.30 dark brown. Antenna dark brown (61,51,38), whitish stripe (241,229,195) composed by fl5 apical 0.60, fl6–12, and fl13 dorsal 0.50. Pronotum black. Propleuron dark brown. Mesoscutum black, median lobe centrally with whitish (233,222,178) spot. Tegula whitish and subalar ridge whitish. Epicnemium with dark brown and orange areas. Mesopleuron bright orange, becoming dark anteriorly; hypoepimeron, mesepimeron, metapleuron and dorsal division of metapleuron bright orange. Scutellum mostly covered by a bilobed whitish (233,222,178) area, basally black. Postscutellum whitish. Propodeum bright orange. Fore leg: coxa bright orange; trochanter, trochantellus and femur light orange (221,166,64); tibia and t1–3 light yellow (228,205,113); t4–5 dark brown. Mid leg: coxa bright orange; trochanter, trochantellus and femur bright orange (225,146,42); tibia light yellow; t1 yellowish brown (170,125,81), its apex and t2–5 brown. Hind leg: coxa, trochanter and trochantellus bright orange (202,115,42); femur reddish brown (162,75,16); tibia light orange, apical 0.25 dark brown; t1 whitish (241,229,195), basal 0.35 dark brown; t2–4 whitish; t5 dark brown. Metasoma striped, dark brown and whitish (233,222,178). T1 basally bright orange, centrally dark brown, posterior margin with whitish stripe (233,222,178). T2–8 bicolored, dark brown with posterior 0.20–0.30 whitish. S2–4 anteriorly dark brown, posteriorly whitish.

Morphometry. See Table 1, Figs 20–21 , and Appendix.

Male. Unknown.

Comments. Structure. Most similar to F. yucatanica , but differs by having the following features: HW1C 1.19 (vs. 1.76); OST 1.70 (vs. 1.37); T2WW 1.91 (vs. 1.75); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); fore wing crossvein 1m-cu basally bent, otherwise straight (vs. more or less uniformly curved); notaulus reaching about 0.54 of mesoscutum length (vs. 0.32); fore wing crossvein 3r-m nebulous (vs. spectral); MLW 1.54 (vs. 1.44); T1WW 2.22 (vs. 1.92); T1LW 2.33 (vs. 2.54); SLW 2.21 (vs. 1.88).

Also similar to F. panamensis , but differs by having the following features: Sternaulus strongly sinuous (vs. weakly sinuous); OST 1.70 (vs. 1.58–1.59); HW1C 1.19 (vs. 1.30–1.49); T1LW 2.33 (vs. 2.67–2.85); fore wing crossvein 1m-cu basally bent, otherwise straight (vs. more or less uniformly curved); MSM 0.25 (vs. 0.33–0.39); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); notaulus reaching about 0.54 (vs. 0.30–0.47) of mesoscutum length; fore wing crossvein 3r-m nebulous (vs. spectral); epomia short, ending far from dorsal margin of pronotum (vs. moderately long, ending near dorsal margin of pronotum).

Also similar to F. exelysae , from which it differs by having the following features: OST 1.70 (vs. 1.43–1.54); sternaulus strongly sinuous (vs. weakly sinuous); MSM 0.25 (vs. 0.29–0.38); area behind anterior transverse carina transversally strigate,arched posteriorwards(vs. transversally strigate,generally weak,antero-laterally faint);fore wing crossvein1m-cu basally bent, otherwise straight(vs. more or less uniformly curved); median depression of T1elongate, very shallow (vs. drop-shaped, distinct, large); fore wing crossvein 3r-m nebulous (vs. spectral); metapleuron densely punctate-rugulose (vs. densely punctate); T2–8 from distinctly coriarious T2 to smooth T8 (vs. faintly coriarious).

Color pattern. Closest to F. pichincha , from which it differs by having pronotum collar black (vs. yellow); propleuron dark brown (vs. light orange); and mandible apical 0.30 dark brown (vs. apical 0.65 blackish).

Etymology. The specific epithet is a literal reference to the ecological reserve Sacha Lodge, where the holotype was collected.

Distribution records. Ecuador.

Material examined. Two specimens, 2♀♀. Holotype: ECUADOR, 1♀, Sucumbios,?, Napo River, Sacha Lodge , 270 m, 0.5°S, 76.5°W, 1994-9-20 /30, P.Hibbs, Malaise, FAS 4626 ,?, CNCI • Paratype: COLOMBIA, 1♀, Putumayo, 2?, PNN La Paya Cabaña La Paya , 330 m, 0˚2′S, 75˚12′W, 2001- 12/12-5/25, E.Lozano, Malaise, 2?, IAVH .

Figures

FIGURE 12. Fortipalpa sacha sp. nov. Holotype female. Habitus.

FIGURE 13. Fortipalpa sacha sp. nov. Holotype female. A, Dorsal view. B, Head, frontal. C, Propodeum, dorsal. D, Ovipositor tip, left side.

FIGURE 19. All known distribution records for Fortipalpa spp. (first map) and individual records for each species. Taxa presented in alphabetical order.

FIGURE 20. All results for morphometric ratios calculated for specimens of Fortipalpa spp., shown comparatively. On each plot, the species are sorted on the Y-axis from highest (top) to lowest (bottom) average. Graphs organized by alphabetical order of ratios.

FIGURE 21. Continuation of Figure 20.