Borboroides, Malloch, 1925

McAlpine, DK, 2007, Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae), with Reconsideration of the Status of Families Heleomyzidae and Sphaeroceridae, and Descriptions of Femoral Gland-baskets, Records of the Australian Museum 59, pp. 143-219 : 158-162

publication ID

2201-4349

persistent identifier

https://treatment.plazi.org/id/03FA0240-6A4A-653A-3660-4D3BB809D57D

treatment provided by

Felipe

scientific name

Borboroides
status

 

Key to species of Borboroides View in CoL

1 Face almost entirely bare and glossy; arista largely white, yellow basally; dorsocentral bristles 1+2; mesoscutum without setulae ( perkinsi View in CoL group) ...................................................................................................... perkinsi View in CoL (p. 176)

—— Face entirely pruinescent; arista brown to blackish; dorsocentral bristles 0+1, 0+2, or 0+3; mesoscutum setulose, at least on central part ................................................................................................................................ 2

2 Mesopleuron (anepisternite) without setulae; mid tibia with large posterior (as distinct from posteroventral) subapical spur; vein 5 almost straight where it borders on discal cell ( Fig. 38); length of discal cell at least 3× distance of its posterodistal angle from margin measured in direction of distal section of vein 5 ( stewarti View in CoL group) .................................................................................................. 3

—— Mesopleuron setulose ( Fig. 102); mid tibia usually without posterior subapical spur; vein 5 usually distinctly curved outwards where it borders on discal cell ( Figs 98, 104); length of discal cell much less than 3× distance of its posterodistal angle from margin ............................................. 15

3 Females .......................................................................................................................................... 4

—— Males .............................................................................................................................................. 9

4 Abdominal segment 5 reduced in size, forming part of retractile postabdomen ( Figs 36, 37), with relatively small tergite; mid tibia always without ventral bristle other than subapical spur ............................................................... 5

—— Segment 5 forming obvious part of ovoid preabdomen, its tergite almost as long as tergite 4; mid tibial armature variable ............................................................... 6

5 Anterior fronto-orbital bristle proclinate; mesopleuron thinly pruinescent, with glossy bare zone restricted to anteroventral quarter; mesoscutum tawny-yellow with median brown stripe narrowed anteriorly and sublateral postsutural brown mark; abdominal tergite 4 not margined; tergite 5 almost parallel- sided ( Fig. 36) ......................................................................................................... musica View in CoL (p. 164)

—— Anterior fronto-orbital bristle eclinate; mesopleuron with large glossy bare central zone, pruinescent posteriorly and anterodorsally; mesoscutum much more extensively brown to black, but partly fading in old specimens; abdominal tergite 4 margined, i.e. with groove parallel to lateral margin setting off convex marginal ridge; tergite 5 almost semicircular ( Fig. 37) ........................................................ donaldi View in CoL (p. 173)

6 Mesoscutum tawny-yellow, usually with median brown stripe and sublateral brown mark; mid tibia without ventral bristles except near apex ................................................................................................................ stewarti View in CoL (p. 162)

—— Mesoscutum with more extensive black to dark grey coloration; mid tibia usually with one or more ventral bristles near or before mid-length ...................................................................................................................................... 7

7 Prelabrum and lower part of occipital region brown to black; mid tibia with a series of 2 to 4 short ventral bristles in addition to subapical ventral spur; tergite 5 extensively dull, pruinescent, particularly on broad median zone and on lateral margins .................................... danielsi View in CoL (p. 167)

—— Prelabrum and lower half of occipital region yellow or tawnyyellow; mid tibia generally with one long ventral bristle in addition to subapical spur; tergite 5 largely glossy, pruinescent mainly on anterior and posterior margins ........................................................................... 8 8 Anterior fronto-orbital bristle not more than half as long as posterior one, slightly reclinate; hind femur with only one anterodorsal bristle, in addition to any small subapical one; abdominal sternite 5 less than 1.4× as long as sternite 4, entirely densely pruinescent ............................................................................................................. tonnoiri View in CoL (p. 171)

—— Anterior fronto-orbital bristle more than half as long as posterior one, slightly proclinate; hind femur usually with two longitudinally aligned anterodorsal bristles in addition to small subapical one; abdominal sternite 5 more than 1.5× as long as sternite 4, much of posterior third only thinly pruinescent and shining ...................................................................................... lindsayae View in CoL (p. 169)

9 Mid tibia with long ventral mollisetae, many of them more than twice as long as tibial diameter ( Fig. 52) ..................................................................................... 10

—— Mid tibia with any mollisetae short and inconspicuous ( Fig. 28) ................................................ 11

10 Anterior fronto-orbital bristle reclinate and slightly sloped outwards, up to c. half as long as posterior one; intradorsocentral setulae in pair of paramedian series, with very few setulae between these and line of dorsocentral bristles; cercus with dense, conspicuous armature of bristles and mollisetae ( Fig. 51) .................................... tonnoiri View in CoL (p. 171)

—— Anterior fronto-orbital bristle eclinate and slightly sloped forwards, c. two-thirds as long as posterior one; intradorsocentral setulae in two well-developed pairs of series; cercus with relatively sparse, inconspicuous setulae and mollisetae ( Fig. 50) .................................................. lindsayae View in CoL (p. 169)

11 Hind tibia strongly curved and attenuated basally ( Fig. 30); cercus small, with setulae short in single marginal series plus pair on dorsobasal prominence; mesoscutum tawny-yellow, with median brown stripe and sometimes pair of brown sublateral marks ...................................................................................................................... stewarti View in CoL (p. 162)

—— Hind tibia almost straight; cercus generally larger, with different arrangement of setulae; thorax variably marked, sometimes much darker than above ............................................................................................................... 12

12 Anterior fronto-orbital bristle proclinate; cercus with conspicuous brush of long mollisetae; mesoscutum tawny-yellow with median and sublateral brown stripes ....................................................................... musica View in CoL (p. 164)

—— Anterior fronto-orbital not proclinate; mollisetae on cercus less developed; mesoscutum of different coloration ........................................................................... 13

13 Mesopleuron with large central bare glossy zone (pruinescent towards dorsal and posterior margins); middle dorsocentral bristle slightly shorter than anterior one (rarely vestigial) ................................................ donaldi View in CoL (p. 173)

—— Mesopleuron densely pruinescent except near anteroventral angle; middle dorsocentral bristle about as long as anterior one ............................................................ 14

14 Entire occiput and prelabrum, and at least part of face and fore coxa brown to blackish; cheek less than half as high as eye; surstylus broad, less than twice as long as wide ..................................................... danielsi View in CoL (p. 167)

—— Lower occiput, prelabrum, face and fore coxa tawny-yellow; cheek c. half as high as eye; surstylus at least three times as long as width at mid-length ........................................................................ probable variant of lindsayae View in CoL (p. 169)

15 Anal cell open distally ( Fig. 92); arista more than 1.5× as long as maximum diameter of eye; mid femur with only one well differentiated anterior bristle ( Fig. 91); male: epandrium less than 0.13 mm long ( parva View in CoL group) ............................................................................. parva View in CoL (p. 187)

—— Anal cell closed ( Fig. 76); arista usually less than 1.5× as long as maximum diameter of eye; mid femur with a series of well differentiated anterior bristles; male: epandrium more than 0.22 mm long ....................................................................................................................... 16

16 Fronto-orbital bristles two large pairs, anterior one long, directed outwards ( Fig. 109); abdominal sternite 1 well sclerotized on whole width; distal section of vein 6 usually less than twice as long as anal crossvein ( atra View in CoL group) ......................................................................................... 17

—— Fronto-orbital bristles one large pair, sometimes also a much smaller reclinate anterior pair ( Figs 65, 80); abdominal sternite 1 reduced, sclerotized mainly at sides; distal section of vein 6 at least twice as long as anal crossvein ..................................................................................... 27

17 Dorsocentral bristles three well-developed pairs; prescutellar acrostichal bristle developed; pleural region with narrow zone of dense grey pruinescence covering entire margin of fore coxal foramen; subapical anteroventral spur of hind tibia much longer than tibial diameter ( Fig. 103); wing veins mostly yellowish ............................................................................................................ gorodkovi View in CoL (p. 193)

—— Dorsocentral bristles usually one or two pairs; prescutellar acrostichal bristle undifferentiated or almost so; pleural region bordering fore coxal foramen shining, at least on upper part, with little pruinescence mainly towards lower part of foramen; subapical anteroventral spur of hind tibia generally not distinctly longer than tibial diameter, sometimes minute; wing veins brown ........................................................ 18

18 Dorsocentral bristles two, anterior one nearly as long as posterior one; section of costa on second costal cell with numerous setulae as long as or longer than width of cell; anal crossvein very oblique on large part, so as to make posterodistal angle of anal cell very obtuse; subapical anteroventral spur of hind tibia absent; male: cerci complex, fused basally, but forming pair of long, free distal lobes, the whole complex longer than epandrium ( Fig. 99) ............................................. menura View in CoL (p. 190)

—— Only one long (posterior) dorsocentral bristle present, any others quite small; section of costa on second costal cell with setulae all shorter than width of cell; anal crossvein almost transverse or slightly sloping distad posteriorly; subapical anteroventral spur of hind tibia usually distinct (vestigial in B. merzi View in CoL ); male: cercus much shorter than epandrium ................................................................................. 19

19 Postfrons, including part in front of anterior ocellus, dull-pruinescent ( Fig. 109); cheek c. half as high as eye; mesopleuron with very extensive posterior grey-pruinescent zone; male: protandrium stout, as broad as epandrium ........................................................................................................ 20

—— Postfrons with distinct, sometimes small, ovate to subtriangular glossy zone in front of anterior ocellus ( Figs. 121, 140), or more extensively glossy; height of cheek usually less than half height of eye; mesopleuron with less extensive posterior pruinescent zone; male: protandrium usually more slender than epandrium, at least posteriorly from mid-length ............................................................................................. 21

20 Postfrons with black ground colour, sometimes tending dark brown anteriorly; cheek entirely black; palpus dark grey to black; male: fore tibia with anteroventral concavity near mid-length; surstylus ( Figs 117, 118) basally transversely broadened, curved before mid-length, with slender almost straight distal section directed posteroventrally, without preapical gibbosity ..................................................... corynetes View in CoL (p. 198)

—— Postfrons partly black, with anterior margin yellow to orange- tawny, more conspicuously so in male; cheek yellowish anteriorly, at least in male; palpus yellow to yellow-brown; male: fore tibia without anteroventral concavity; surstylus ( Figs 114, 115) somewhat variable with distal section not posteriorly inclined, typically with posterior preapical gibbosity .............................................................. shippi View in CoL (p. 195) 21 Capitellum of halter grey-brown; male: surstylus with large anterior, outwardly inclined rounded tubercle at base ( Fig. 149); cercus dorsally fused to epandrium ............................................................................. atra View in CoL (p. 211)

—— Capitellum yellow or whitish; male: surstylus without such outwardly inclined tubercle; cercus articulated with epandrium ...................................................... 22

22 Postfrons with small median glossy zone, much narrower than lateral subshining (non-pruinescent) zone ( Fig. 121); cheek more than one third as high as eye; male: segment 8 slender throughout; epandrium very broad; cerci clavate, narrow and widely separated basally ( Fig. 123) ....................................................................... petiolus View in CoL (p. 200)

—— Postfrons with larger median glossy zone, c. as wide as or wider than lateral subshining zone ( Figs 125, 130); cheek usually one third as high as eye or less; male: abdominal segment 8 widened anteriorly or moderately broad throughout; epandrium less than twice as broad as segment 8; cerci variable but not widely separated and clavate ....................................................................................................... 23

23 Postfrons very extensively shining, with little pruinescence even on anterior half; male: epandrium on each side with two large anteroventral bristles, each at least one third as long as epandrium; surstylus distally on inner surface armed with complex, fascicle-like spines; cercus rod-like, long and slender ( Fig. 154); habitat southwestern Australia............................................................................ woodhilli View in CoL (p. 212)

—— Postfrons pruinescent on much of anterior half; male: epandrium without such large bristles; surstylus with simple spines and/or setulae only; cercus relatively broad, not rod-like; habitat eastern Australia (including Tasmania) ................................................................................................... 24

24 Postfrons with pair of sublateral shining zones resembling median glossy zone, but not fused with it; subapical spur of hind tibia nearly as long as greatest diameter of tibia; male: surstylus only slightly widened basally, almost straight; cercus plate-like, with very long fringe of mollisetae ( Fig. 129); female: tergite 5 with median grey-pruinescent zone on full length .......................................................... fimbria View in CoL (p. 203)

—— Postfrons either without markedly shining sublateral zones extending in front of level of anterior ocellus, or median and sublateral zones fused to form broad shining zone; subapical spur of hind tibia generally much shorter than greatest diameter of tibia or indistinguishable; male: surstylus very broad basally, curved; cercus with short setulae only; female (unknown in B. acumen View in CoL ): tergite 5 pruinescent anteriorly, but without such complete median pruinescent zone ...................................................................................... 25

25 Male: protandrium scarcely narrowed posteriorly; epandrium ( Fig. 142) broadly attached to protandrium, dorsobasally prominent and, when in anteriorly flexed position, projecting posteriorly beyond end of protandrium; cercus small, transverse; female: tergite 6 undivided; cercus slightly bulbous apically ............................................................... merzi View in CoL (p. 206)

—— Male: protandrium narrowed posteriorly; epandrium shortly attenuated at base and not strongly projecting; cercus large, transversely compressed; female (unknown in B. acumen View in CoL ): tergite 6 divided by median desclerotization; cercus not thickened apically ............................................. 26

26 Postfrons with subtriangular, median glossy zone, but without markedly shining non-pruinescent lateral zones extending in front of level of anterior ocellus; male: surstylus ( Fig. 133) with large, broadly rounded anterior basal dilation, its slender apex directed anteriorly; cercus ovate, obtuse ................................................................ bulberti View in CoL (p. 205)

—— Postfrons with lateral glossy zones fused with median zone to form broad glossy zone extending in front of anterior ocellus; male: surstylus ( Fig. 146) broadly spatulate, without such rounded anterior basal expansion; cercus broad, shortly narrowed to subacute apex ..................................................................................................... acumen View in CoL (p. 209)

27 Postfrons in front of level of anterior ocellus entirely dull, with fingerprint sculpture and / or pruinescence; sternopleuron almost entirely densely pruinescent ( staniochi View in CoL group) ............................................................................ 28

—— Postfrons with extensive smooth, glossy zone in front of anterior ocellus; sternopleuron shining and bare, at least in part ( helenae View in CoL group) ............................................................................................................................ 29

28 Mesopleuron very extensively pruinescent, glossy only on small zone near anterior margin ( Fig. 66); cheek yellow in large part (at least in male); mid tibia in both sexes with well-developed subapical posterior spur and ventral spur; distal section of vein 6 extending three quarters of distance from anal crossvein to posterior wing margin; male: cercus ( Fig. 67) small and narrow ................................ dayi View in CoL (p. 178)

—— Mesopleuron glossy centrally and anteriorly ( Fig. 74); cheek entirely black to dark brown; mid tibia in both sexes with at most very small subapical posterior spur, with subapical ventral spur in female only; distal section of vein 6 extending no more than two thirds distance to margin; male: cercus ( Figs 77, 79) broad ................................ staniochi View in CoL (p. 180)

29 Mesopleuron largely glossy black, with small pruinescent zone on dorsal margin only ( Fig. 81); fore femur with at least three long posteroventral bristles, but no anteroventral comb; fore tarsus entirely brown; capitellum of halter yellowish brown to dark brown ......................................................................................................... helenae View in CoL (p. 183)

—— Mesopleuron extensively pruinescent, on both dorsal and posterior margins; fore femur with posteroventral bristles little developed, but, in male only, with comb of stout, moderately short anteroventral bristles; fore tarsus bicoloured, segment 1 dark brown, segments 2 to 5 pale yellowish; capitellum of halter creamy-white ....................................................................................................... doreenae View in CoL (p. 186)

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Heleomyzidae

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