Amplirhagada basilica, Köhler, 2010

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 264-266

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
persistent identifier	https://treatment.plazi.org/id/03F987C5-FFBE-F76D-01D0-15CB4C35C9A7
treatment provided by	Felipe
scientific name	Amplirhagada basilica
status	sp. nov.

Treatment

Amplirhagada basilica View in CoL n.sp.

Type locality ( Fig. 1 View Figure 1 ). Western Australia, Kimberley, Bonaparte Archipelago, Augustus Island , southern section, Brecknock Harbour , 15°23'53"S 124°036'03"E; KIS 2-51. Weathered south facing sandstone scree in deep gorge, under rocks (leg. R. Teale, 1 June 2008) GoogleMaps .

Type material. Holotype WAM S34623 View Materials (Pl. 2.11) . Paratypes AMS C463720 (4 preserved specimens, same as holotype), WAM S37004 View Materials (11 preserved specimens, same as holotype), AMS C463721 (3 shells, 15°22'54"S 124°35'24"E), WAM S37414 View Materials (4 shells, 15°22'54"S 124°35'24"E) GoogleMaps .

Additional, non-type material. WAM S37000 View Materials –3, WAM S37413 View Materials (Augustus Island).

Etymology. From basilica (Latin = royal, splendid, magnificent), referring to its large shell. Amplirhagada regia is among the largest species of the genus.

Sealing strategy. Free sealer. Shell ( Fig. 73A–C View Figure 73 , Pl. 2.11). Semi-globose, with medium high spire; solid to thick. Periphery slightly angulate; upper and basal sectors of whorls rounded. Umbilicus open, forming a chink to narrowly winding opening, 80–95 percent concealed by columellar reflection. Background colour blends from cream into brownish (upper sector of lower whorls darker); sub-sutural band diffuse, light brownish; mid-whorl band diffuse, thin, brownish, most pronounced, visible on last whorls only, blurring towards tip of shell; ventral colour horn; outer lip colour differs from shell, brownish purple; inner lip colour pinkish. Protoconch 2.4 mm in diameter, comprising 1.7 whorls, smooth. Teleoconch smooth except of indistinct growth lines. Angle of aperture 45°; outer lip rounded, moderate to thick, expanded, slightly reflected, basal node of lip weak, palatal node absent. Parietal wall of inner lip absent.

stimulatory pilaster absent. Two smooth, narrow longitudinal pilasters comprise entire length of penial chamber; two additional smooth, narrow pilasters develop at base of penial chamber. Vas deferens rather thick, entering penial sheath apically. Vagina rather short, posteriorly inflated; inner vaginal wall densely covered by tiny pustules and smooth longitudinal pilasters. Spermatheca long, extending over base of spermoviduct. Spermathecal duct moderately

Pallial morphology. Pallial cavity moderately deep, extending 3 ⁄ 4 whorl; mantle pigmentation mottled, dark grey. Kidney extending about half of pallial cavity.

Genital morphology ( Figs. 74–75 View Figure 74 ). Penis straight, shorter than anterior part of oviduct, very thin. Vas deferens forms simple loop before entering penis. Penial retractor muscle of same length as penis. Penial sheath evenly thick. Penial verge comprising 1 ⁄ 8– 1 ⁄ 4 of penial chamber, slender to spatulate, with rounded tip. Penial wall pustules absent. Main wide, internally with pustules. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with thin wall. Free oviduct comprising half of anterior part of oviduct or more; more or less straight. Spermoviduct much longer than anterior part of oviduct. Talon embedded in albumen gland anteriorly.

Radular morphology ( Fig. 73E–F View Figure 73 ). Rectangular. Tooth formula C+14+4+22. In average 119±19.0 rows of teeth, 28.0±0.4 rows per mm (n = 2). Central teeth with bluntly pointedovate mesocone, shorter than base of teeth; ectocones vestigial. Lateral teeth with bluntly pointed, ovate mesocones, shorter than base of teeth; ectocones and endocones vestigial. Marginal teeth with broadly rounded mesocones; ectocones pointed, much smaller and narrower than mesocones; endocones vestigial.

Comparative remarks. One of the few species with large, dome-shaped shells. Differs from A. tricenaria and A. regia by dark outer lip. Purplish colour of outer lip similar to A. sphaeroidea . Absence of penial wall pustules is characteristic for this species.