Meoneura joedaltoni, Stuke & Barták, 2019

Stuke, Jens-Hermann & Barták, Miroslav, 2019, Records of Carnidae from the collection of Miroslav Barták (Diptera: Carnidae), with the description of five new species, Zootaxa 4567 (2), pp. 326-346: 338-340

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Meoneura joedaltoni

spec. nov.

Meoneura joedaltoni   spec. nov.

( Figs. 15–18 View FIGURES 15–18 )

Holotype ♂: (1) „I: Amatrice-5kmS, pi- / ne wood nr.pasture / 42.39N / 13.20E 1700 m / Barták 3.viii.1988 “; (2) „ Holotypus / Meoneura joedaltoni   / spec. nov. ♂ / det. Stuke 2018“. GoogleMaps  

The holotype is deposited in the collection of the Czech University of Life Sciences, Czech Republic, Prague (CULSP). The specimen is glued on a card. Otherwise the specimen is in good condition. The posterior part of abdomen is dissected, macerated and stored in a glycerine microvial pinned underneath the specimen.

Description of holotype (male). Length about 1.5 mm. Wing length 1.2 mm.

Head height 0.4 mm. Face, anterior half of frons and gena below the eye orange brown. Posterior half of frons brown with frontal triangle black. Occiput and main part of gena black. Antenna brown. Arista without pubescence. Maximum eye length: maximum eye height = 1.0. Posteroventral margin of gena closest to eye margin: maximum eye height = 0.6. Frons slightly microtomentose, slightly microtomentose. Frontal triangle distinct, reaching about 1/3 distance from anterior ocellus to frontal margin. Face microtomentose. Carina broad, almost as broad as fore tibia. Postcranium microtomentose. Prementum and palpus cannot be examined at the holotype due to the preparation. 1 distinct ocellar seta; supralunular almost parallel; 4 fronto-orbital setae (2 anterior mesoclinate, 2 posterior lateroclinate), the posterior mesoclinate seta short, only 1/3 of the length of the anterior mesoclinate seta. Perhaps this short seta is an additional anterior mesoclinate seta and another larger mesoclinate seta is broken (on both sides of the head). 2 vertical setae; 2 mesoclinate postorbital setae; postocellar setae slightly divergent; 1 strong vibrissal seta; supravibrissal setae incomplete in the holotype; 1 strong and 2 slightly smaller genal setae.

Scutum microtomentose, covered with short, semiadpressed black setulae. Scutellum microtomentose. Pleurae—as far as can be seen in the type—microtomentose. Scutum with 1 long posterior dorsocentral seta only. 2 postpronotal setae; 1 praesutural seta; 2 notopleural setae; 1 supraalar seta; 2 postalar setae; 1 praescutellar seta; 1 apical and 1 lateral scutellar seta. Setae on katepisternum and anepisternum cannot be examined in holotype. 1 dorsal seta and 1 ventral seta on katepisternum. Costa with no obvious setae beyond radial vein R 1. Wing hyaline, veins light yellow to yellow brown. R 4+5 slightly curved to apex of wing. Haltere brown, base of haltere brown. Legs black to brown. Fore femur basally with 2 and apically with 2 strong posteroventral setae. Hind femur apically with 1 strong anteroventral seta. All coxae with single inconspicuous black setae. Hind metatarsus ventrally with dense yellow golden setae. Length metatarsus 2: length tibia 2 = 0.5.

Tergites with no obvious depressions or tufts of setulae. Abdominal pleura with scattered setae on abdominal segments 3–5. Abdominal segments 1–5 narrow, width of tergite 3: length of tergite 3 = 3.1. Tergites 2–5 each with 1 distinct lateral seta at the posterior margin, and tergite 5 with 2 setae at the posterior margin. Sternite 5 wider than long; the posterior margin slightly concave ( Fig. 18 View FIGURES 15–18 ). Sternite 5 with scattered setae in the posterior half. Midventral tergite 7 distinct. Protandrium small, shorter than epandrium and about 0.4 of length of tergite 5. In the holotype the protandrium is almost completely hidden by tergite 5. Epandrium with four outstanding lateral setae—the basal distinctly longer than the three other—and one smaller setae ( Fig. 15 View FIGURES 15–18 ). Maximum length dorsally in the middle of epandrium: maximum width of epandrium = 0.4. Cerci indistinct, not projecting, and no setulae recognized. Subepandrial plate distinctly sclerotised. No setula recognized on subepandrial plate. No proceeding of hypoproct. No tooth on subepandrial plate. Surstylus as in Fig. 16 View FIGURES 15–18 : large, parallel sided, anteriorly rounded. Surstylus with few setulae only. No lamella. Postgonite as in Fig. 17 View FIGURES 15–18 : distinctly sclerotised, broad, hardly elongated, dorsoapically pointed. Distiphallus small, inconspicuous, curved, longer than epandrium and surstylus, covered with dense light brown setulae. Basiphallus obviously sclerotised, long, narrow, at its base widened.

Diagnosis. An exciting character of M. joedaltoni   is the completely brown halter that is otherwise only known from European Meoneura elongella Zetterstedt, 1838   , Nearctic M. nigrifrons Malloch, 1915   and Nearctic M. wirthi Sabrosky, 1959   . While the two Nearctic species have completely different structures at the postabdomen (cf. Sabrosky 1959) the European M. elongella   is only known from female specimens. The most obvious character to distinguish the holotype of M. joedaltoni   and a female of M. elongella   from Norway is the more yellow coloured head (face, gena at eye margin and about half of the frons) of M. joedaltoni   . This character is important in other Meoneura   species, for example to distinguish the closely related species of the M. flavifacies   -group. It is described in Zetterstedt´s description of M. elongella   . Another obvious character is the higher face between antennal groove and upper mouth opening (higher than fore tibia diameter) in M. joedaltoni   .

In regard to the postabdomen, Meoneura joedaltoni   belongs to the Meoneura palaestinensis   -group sensu Stuke & Freidberg (2017) that is characterised by a broad and anteriorly rounded surstylus and the lack of an adjected lamella ( Fig. 16 View FIGURES 15–18 ). M. joedaltoni   is easily identified by this unique combination of characters: (1) face and anterior half of frons orange yellow; (2) carina about as broad as fore tibia; (3) scutum and scutellum distinctly microtomentose; (4) 1 dorsocentral only; (5) haltere completely brown; (6) epandrium with 1 outstanding and 3 distinct setae ( Fig. 15 View FIGURES 15–18 ); (7) cercus indistinct and without recognized setae; (8) surstylus as in Fig. 16 View FIGURES 15–18 : large, almost as long as epandrium, parallel sided, not narrowing basally, apically rounded, not tapering and not hyaline apically; (9) hypoproct without any proceeding; (10) sternite 5 slightly wider than long, hind margin slightly concave ( Fig. 18 View FIGURES 15–18 ). A key of the Meoneura palaestinensis   -group was published by Stuke & Freidberg (2017). M. joedaltoni   will run to the last couplet 4 together with M grimmorum Stuke & Freidberg, 2017   and M. palaestinensis Hennig, 1937   , but will not be assigned to one of these species due to the very different shape of the surstylus and the combination of characters mentioned above.

Etymology. The species is named after Joe Dalton, the smallest of the four Dalton brothers, who appear in the Lucky Luke comic book series of Maurice de Bevere (Morris) and René Goscinny. Joe is the smallest of the four Dalton brothers as Meoneura joedaltoni   is one of the smallest Acalyptratae.

Distribution. Meoneura joedaltoni   has been caught only at the locus typicus, a pine wood 5 km south of Amatrice (Latium, Italy). The coordinates of the label don´t fit exactly to this description but are about 4 km ssw of Amatrice.

Meoneura lacteipennis   ( FALLÉN, 1823)

Material: CZECH REPUBLIC: 1♂, 22.v.1995, Vráž u Písku , near pond [49.23°N 14.08°E], 400 m, leg. M. Barták, coll. CULSP GoogleMaps   ; TURKEY: 1♂, 17.–22.v.2011, Muğla, University campus, 700 m [37°09’42’’N 28°22’21’’E], 700 m, pan trap +sweeping, leg. M. Barták & Š. Kubik, coll. CULSP GoogleMaps   .

Diagnosis: Collin (1930).

Meoneura lamellata   COLLIN, 1930

Material: CZECH REPUBLIC: 1♂,–7.vii.2015, Praha, Troja [50°07’15’’N 14°23’53’’E], 184 m, emergence trap, pig caracas, exp. 9.xii.2014, leg. M. Barták, coll. PJHS GoogleMaps   ; 1♂, 27.v.–, dito, coll. CULSP   ; 1♂, 22.–29.v.2012, Praha, Troja [50°07’15’’N 14°23’53’’E], 184m, emergence trap baited with pig caracas, leg. M. Barták, coll. CULSP GoogleMaps   ; 1♂,–3.vii.2012, dito.

Diagnosis: Collin (1930).

M. lamellata   is new for the Czech Republic.