Baeocera inoptata, Löbl, 2018

Löbl, Ivan, 2018, A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae), Acta Entomologica Musei Nationalis Pragae 58 (1), pp. 151-165 : 152

Baeocera inoptata sp. nov.

( Figs 1–5 )

Type locality. Indonesia, South Sulawesi, Gowa District, 6 km E of Malino, Gunung Bawakaraeng [Mt.] Area, near Lembanna Base camp, ca. 05°15.4 ′ S, 119°54.5 ′ E.

Type material. HOLOTYPE: ( NMPC), INDONESIA, S SULAWESI: Gowa Distr. / 6 km E of Malino, Gn. Bawakaraeng Area / border of gardens and mixed forest / (dominant Pinus) nr Lembanna Base camp / 05°15.4 ′ S, 119°54.5 ′ E, 1520 m / J.Hájek & J.Šumpich leg., 11-13.ii.2015. PARATYPE: 1 ♀ ( MHNG), with the same data as the holotype.

Description. Length 2.03 mm, width 1.42 mm. Body convex dorsally; black, elytral apices and apical abdominal segments blackish-brown, mouth-parts ochreous, femora dark reddish-brown near bases and apices, blackish in middle, tibiae dark reddish-brown, tarsi ochreous, lighter than tibiae, antennomeres I to III ochreous, following antennomeres darkened, brown. Eyes large. Length/width ratios of antennomeres as: III 20/10: IV 20/8: V 32/8: VI 35/15: VII 55/20: VIII 43/16: IX 57/20: X 55/ 23 ( Fig. 1 ; antennae of the holotype broken off, paratype with one antenna only, lacking apical antennomere). Lateral contours of pronotum and elytra separately rounded. Pronotum with lateral margin carinae concealed near angles, discal punctation very fine. Tip of scutellum exposed. Elytra weakly narrowed apically, middle section of lateral margins straight, lateral margin carinae exposed in dorsal view, sutural striae curved at base and extending laterally to form basal striae joined with lateral striae, adsutural areas flat. Elytral punctation on inner halves of disc somewhat coarser than pronotal punctation and finer than on outer discal halves. Hind wings fully developed. Hypomera very finely punctate. Mesoventrite with very shallow mesal stria, lacking large marginal punctures. Mesepimera large, slightly narrowed mesally, about 5 times as long as wide and 6 times as long as intervals to mesocoxae. Median part of metaventrite convex, smooth in middle, with several coarse punctures near metacoxal process. Lateral parts of metaventrite appearing impunctate. Metacoxal process flat, submesocoxal areas 0.03 mm, with marginal punctures coarse, not elongate, extended to inner margins of mesepimera. Metanepisterna flat, about 0.13 mm wide, with suture deep, slightly sinuate. Abdominal ventrite I not microsculptured, with coarse, not elongate basal punctures, lacking basal rugosity; following ventrites distinctly punctate; ventrite VI with punctulate microsculpture. Tibiae straight.

Male characters. Protarsomeres I–III distinctly widened, narrower than apices of protibiae. Mesotarsomeres I and II slightly widened. Aedeagus ( Figs 2–4 ) 0.63 mm long, strongly sclerotized, asymmetrical. Apical process of median lobe short, robust, overlapped by basal bulb. Internal sac with complex sclerites and two apical tufts of spine-like structures. Right parameres simple, bent in middle, in apical half almost evenly wide seen in ventral view, sinuate and widened in apical section in lateral view; left paramere lacking apophyse, wide, expanded ventrally.

Female characters. Protarsomeres not widened. Gonocoxite almost straight, evenly narrow, with subapical pores ( Fig. 5 ).

Differential diagnosis. Only a single species of Baeocera , B. derougemonti Löbl, 1983 , has been described from Sulawesi. This species is a member of the B. lenta species group defined by symmetrical aedeagi with a long ejaculatory duct extruded in resting position. The new species, a member of the B. monstrosa species group, defined by asymmetrical, voluminous aedeagi with complex internal sacs, with the ejaculatory duct not extruded in repos (see LÖBL 1971). The sole known Indonesian member of the B. monstrosa group is the Sumatran B. robertiana Löbl, 1990, a species that differs drastically from B. inoptata sp. nov. by the shape of the parameres: it has the right paramere strongly expanded and lamellar, abruptly narrowed and denticular near the tip. Also notable is the internal sac in B. robertiana lacking apical tufts of spine-like structures. Both species may be easily distinguished by their antennae: the antennomere IV in B. robertiana is about 1.5 times as long as antennomere III and similar to antennomere VI, while antennomeres III and IV are equally long in B. inoptata sp. nov. with III much shorter than antennomere VI. The median lobe and the parameres of the new species are comparatively similar with those of the Philippine B. alticola Löbl, 2012. However, the internal sac bearing robust, elongate sclerites in the latter species is distinctive (see LÖBL 2012b). The “ Baeocera sp1” in OGAWA (2015) is likely B. inoptata sp. nov.

Etymology. The species epithet is a Latin adjective inoptatus, - a, - um, meaning ‘not wanted’.

Distribution. Indonesia, South Sulawesi.

Figures

Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18).