Scaphisoma, Leach, 1815

Löbl, Ivan, 2018, A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae), Acta Entomologica Musei Nationalis Pragae 58 (1), pp. 151-165 : 162-164

Key to Sulawesi species of Scaphisoma

1 Pronotum and elytra both with distinct bicoloured colour pattern. Elytra with sutural striae not curved along pronotal lobe. ............ S. versicoloreum sp. nov.

‒ Pronotum unicoloured, elytra about as unicoloured as pronotum, or with distinctive colour pattern. ........... 2

2 Elytra bicoloured, mostly dark, each with yellowish, well delimited, subapical transverse fascia. ................... ................................................ S. flavolineatum sp. nov.

‒ Elytra either unicoloured or if bicoloured, elytral pattern consist of reddish spots; never with light subapical fasciae. ................................................................... 3

3 Elytra with sutural striae extending along bases to form basal striae. Male protarsomere I strongly enlarged, as wide as or wider than tibial apices. ................ 4

‒ Elytra with sutural striae ending near pronotal lobe, not extended laterad to form basal striae. Male protarsomere I usually narrower than tibial apices. ............ 5

4 Length 2.15 to 2.45 mm. Body dark brown to black, mesepimera about as long as third of intervals between them and mesocoxae. Abdominal ventrite I microsculptured. ...................... S. latitarse Löbl, 2012

‒ Length 1.90 to 2.24 mm long. Body light, reddish- -brown, mesepimera shorter than intervals between them and mesocoxae. Abdominal ventrite I not microsculptured. ................................ S. hulai sp. nov.

5 Elytra with sutural striae shortened, starting behind level of scutellum. .......................................................... 6

‒ Elytra with sutural striae not shortened, starting at elytral bases. ................................................................... 8

6 Length 1.15 to 1.20 mm, pronotum and elytra uniformly reddish-brown. Pronotum with punctation coarser near lateral margins than on centre of disc. ........... ........................................................... S. napu Löbl, 1983

‒ Length 2.20 mm, elytra on apical fourth notably lighter than on remaining surface and pronotum. Pronotal punctation even throughout. .................................. 7

7 Metaventrite with mesal carina. Abdominal ventrite I entirely very finely punctate. .... S. carinatum sp. nov.

‒ Metaventrite lacking mesal carina. Punctation on middle of abdominal ventrite I much coarser than on lateral parts. .......................................... Scaphisoma sp.

8 Elytra with distinct bicolorous pattern. ...................... 9

‒ Elytra entirely light reddish-brown to black, or with narrowly lighter apical margins, or somewhat darkened near apical margins. ............................................. 12

9 Pronotum and elytra dark, blackish-brown to black, elytra each with one or two reddish spots, body length 1.20 to 1.40 mm. ......................................................... 10

– Elytral colour pattern different, body length 1.7 to 2.0 mm. .............................................................................. 11

10 Elytra each with single reddish spot and with reddish apical area. Aedeagus with trifid apical process, internal sac with sclerotized, teeth-like structures. .............. .................... S. obliquemaculatum Motschulsky, 1863

– Elytra each with two reddish spots, lacking light apical area. Aedeagus with apical process simple, internal sac membranous, lacking sclerotized structures. ............... ............................................................ S. bugi Löbl, 1983

11 Elytral punctation very dense and coarse, much coarser than pronotal punctation, puncture diameters in part larger than puncture intervals. Metaventrite with antecoxal puncture row. Aedeagus symmetrical, with apically widened, lobed parameres. ............................... ............................................................ S. palu Löbl, 1983

– Elytral punctation fine and sparse, slightly coarser than that on pronotum; puncture diameters much smaller than puncture intervals. Metaventrite without antecoxal puncture rows. Aedeagus asymmetrical, with parameres not lobed, narrowed apically. .............. ........................................................ S. sumpichi sp. nov.

12 Body length 0.90 mm long. Elytral punctation becoming coarser apically. Abdomen lacking obvious microsculpture. ........................... S. sadang Löbl, 1983

– Body length 1.55 to 1.80 mm. Elytral punctation even. Abdomen microsculptured. ...................................... 13

13 Head and pronotum blackish, elytra with well visible, sharply delimited discal punctures, elytral apices not sexually dimorphic, with apical margins slightly rounded or truncate. Metaventrite with mesal stria, abdominal microsculpture strigulate. .............................. 14

– Pronotum and elytra reddish-brown, elytral punctures hardly visible, not clearly delimited. Elytral apices sexually dimorphic, in male truncate, in female expanded and strongly oblique. Metaventrite lacking mesal stria, abdominal ventrite I not microsculptured, following ventrite with punctulate microsculpture. ............. ............................................................ S. ancora sp. nov.

14 Metaventrite bearing strigulate microsculpture. Aedeagus with internal sac lacking distinct spine-like structures. ......................................... S. ogawai sp. nov.

– Metaventrite lacking microsculpture. Aedeagus with internal sac bearing distinct spine-like structures. ...... .............................................................. S. pellax sp. nov.