Scaphisoma versicoloreum, Löbl, 2018

Löbl, Ivan, 2018, A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae), Acta Entomologica Musei Nationalis Pragae 58 (1), pp. 151-165 : 159-160

Scaphisoma versicoloreum sp. nov.

( Figs 35–39 , 53 )

Type locality. Indonesia, South Sulawesi, Gowa District, 6 km E of Malino, Gunung Bawakaraeng [Mt.] Area, near Lembanna Base camp, ca. 05°15.4 ′ S, 119°54.5 ′ E.

Type material. HOLOTYPE: ( NMPC), INDONESIA, S SULAWESI: Gowa Distr. / 6 km E of Malino, Gn.Bawakaraeng Area / border of gardens and mixed forest / (dominant Pinus) nr Lembanna Base camp / 05°15.4 ′ S, 119°54.5 ′ E, 1520 m / J.Hájek & J.Šumpich leg., 11-13.ii.2015. PARATYPES: 3 2 ♀♀ ( NMPC, MHNG), with the same data as the holotype.

Description. Length 2.20–2.86 mm, width 1.35–1.50 mm. Head ochreous; pronotum dark brown to black, with narrow mesal ochreous band, anterior margin narrowly ochreous and with conspicuous circular ochreous band at each side; prevailing elytral surface ochreous, basal area black and with sharply delimited and irregular margin, triangularly expanded in middle, adsutural area dark brown, centre of disc with rather small black spot narrowly separated from subtriangular black area touching sutural stria, subapical area with small spot ( Fig. 53 ); hypomera ochreous, with dark margins, venter of mesothorax and metathorax blackish, ventrites reddish-brown, becoming gradually lighter apically; femora and tibiae ochreous, mouth-parts, antennomeres I and II, and tarsi yellowish, antennomeres III to XI brown.Antennae long, length/width ratios of antennomeres as: III 15/8: IV 49/7: V 65/7: VI 58/8: VII 65/15: VIII 50/10: IX 62/14: X 52/14: XI 65/14. Pronotum and elytra not microsculptured. Pronotum strongly narrowed anteriad, with regularly rounded lateral margins, lateral margins carinae entirely or almost entirely visible; punctation on disc fine, dense, visible at magnification 20×, consisting of sharply delimited punctures, puncture intervals much larger than puncture diameters, punctation along lateral carinae distinct, conspicuously dense. Tip of scutellum exposed. Elytra strongly narrowed apically, lateral margins almost regularly curved, lateral margin carinae exposed throughout in dorsal view, apices sexually dimorphic, apical margins rounded, inner apical angles situated posterior level of outer angles, apical serration conspicuous, sutural margin not raised, adsutural areas flat, narrow, each with single puncture row. Sutural striae parallel in basal halves, converging apically, starting at level of scutellum, not curved along pronotal lobe. Punctures along lateral margins coarse and dense. Elytral disc with punctures larger than those on pronotum, sharply delimited, puncture intervals mostly about as large to twice as large as puncture diameters. Hind wings fully developed. Hypomera smooth, not microsculptured. Mesepimera slightly shorter than intervals between them and mesocoxae and four times as long as wide. Metaventrite with strigulate microsculpture; centre of metaventrite convex, with two shallow apical impressions, lacking mesal stria, punctation dense and fine, denser in impressions than on remaining surface; apical intercoxal margin truncate. Lateral parts of metaventrite sparsely and very finely punctate, antecoxal puncture rows absent. Submesocoxal lines convex, distinctly punctate; submesocoxal areas 0.04 mm, about as long as fifth to fourth of shortest intervals between them and metacoxae. Metanepisterna flat, hardly narrowed anteriad, with broadly rounded angles. Protibiae straight. Mesotibiae and metatibiae slightly bent. Abdomen with strigulate microsculpture and extremely fine punctation; ventrite I with submetacoxal lines parallel, coarsely punctate; submetacoxal areas each 0.05 mm, as long as fourth of shortest intervals between them and apical margin of ventrite.

Male characters. Protarsomeres I–III strongly widened, I about as wide as apices of protibiae. Mesotibiae thicker and more bent than metatibiae. Mesotarsomeres I and II distinctly widened. Elytra not extended apically, with inner apical angles lying somewhat posterior level of outer angles. Apical margin of abdominal ventrite VI with narrow, 0.05 mm long lobe. Tergite IX with stalk rounded laterally, bearing basal, not clearly delimited darkened areas ( Fig. 38 ). Aedeagus ( Figs 35–37 ) 0.88–0.90 mm long, symmetrical. Apical process long, tapering, inflexed, with slightly sinuate ventral margin in lateral view. Dorsal valve partly sclerotized to form rhomboid plate joined by membranes to proximal part of apical process. Parameres bent at bases and apices, notched ventrally near apices, with sinuate dorsal margin in lateral view, each bearing ventral, membranous apophyse extended apically. Internal sac with pair of distal denticles and complex basal sclerite, joined by membranes bearing very finely striate and minute denticle-like structures.

Female characters. Pro- and mesotarsomeres narrow. Elytra with prominent apical angles, apical margins strongly oblique, inner apical angles situated far posterior outer apical angles. Gonocoxite slightly bent, evenly narrow ( Fig. 39 ).

Differential diagnosis. The colour pattern of the body and the aedeagal characters of this species are unique. The median lobe is similar to that of S. bugi and its allies, and of the Malaysian S. angulatum Löbl, 1975, but the parameres bearing ventral hyaline apophyses are distinctive. An unusual feature are the sexually dimorphic elytra (see under S. ancora sp. nov.).

Etymology. The species epithet is a Latin adjective versicoloreus, - a, - um, meaning varicoloured.

Distribution. Indonesia, South Sulawesi.

Figures

Figs 35–49. 35–39 – Scaphisoma versicoloreum sp. nov.: 35, 36 – aedeagus in dorsal and lateral views; 37 – internal sac of aedeagus; 38 – male tergite 9; 39 – gonocoxite.40–46 – S. carinatum sp. nov.: 40 – male tergite 9; 41 – apical process of aedeagal median lobe in lateral view; 42 – tip of apical process in ventral view; 43, 44 – aedeagus in dorsal and lateral views; 45 – parameres in ventral view; 46 –internal sac of aedeagus. 47–49 – Scaphobaeocera jirkai sp. nov.: 47, 48 – aedeagus in dorsal and lateral views; 49 – gonocoxite. Scale bars: 0.1 mm (Figs 37, 39, 41–42, 46–49); 0.2 mm (Figs 35–36, 43–45); 0.3 mm (Figs 38, 40).

Figs 50–53. Habitus of Scaphisoma. 50, 51 – S. ancora sp. nov. (50 – male, 51 – female); 52 – S. sumpichi sp. nov., male; 53 – S. versicoloreum sp. nov., male.